Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Leptodactylus elenae Heyer, 1978

Class: Amphibia > Order: Anura > Family: Leptodactylidae > Subfamily: Leptodactylinae > Genus: Leptodactylus > Species: Leptodactylus elenae

Leptodactylus elenae Heyer, 1978, Sci. Bull. Nat. Hist. Mus. Los Angeles Co., 29: 45. Holotype: LACM 92096, by original designation. Type locality: "Argentina: Salta, Embarcación".

Common Names

Marbled White-lipped Frog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 81).

Distribution

Gran Chaco of Argentina (Jujuy and Salta), Bolivia (Beni, La Paz, and Santa Cruz), throughout Paraguay, and central Brazil (Mato Grosso and Mato Grosso do Sul); one record for Corrientes, northeastern Argentina. Records for souheastern Peru and controversial (see comment)

Geographic Occurrence

Natural Resident: Argentina, Bolivia, Brazil, Paraguay, Peru

Comment

In the Leptodactylus fuscus group of Heyer, 1978, Sci. Bull. Nat. Hist. Mus. Los Angeles Co., 29: 1–85. Cei, 1987, Monit. Zool. Ital., N.S., Suppl., 21: 223–225, noted that the species reviewed by Cei, 1980, Monit. Zool. Ital., N.S., Monogr., 2: 338–340, under the name Leptodactylus mystaceus was of this species. Márquez, De la Riva, and Bosch, 1995, J. Zool., London, 237: 313–336, reported on vocalization in Bolivia. Reviewed by Heyer and Heyer, 2002, Cat. Am. Amph. Rept., 742: 1–5. Zaracho and Alvarez, 2005, FACENA, 21: 135, provided a record for the Parque Nacional Mburucuy, Corrientes, Argentina. Brusquetti and Lavilla, 2006, Cuad. Herpetol., 20: 13, briefly discussed range in Paraguay. Jansen, Bloch, Schulze, and Pfenninger, 2011, Zool. Scripta, 40: 567–583, suggested on the basis of molecular evidence that at least one unnamed species may exist in Bolivia. Weiler, Núñez, Airaldi, Lavilla, Peris, and Baldo, 2013, Anf. Paraguay: 90, provided a brief account, image, and dot map for Paraguay. In the Leptodactylus fuscus species group, Leptodactylus mystaceus complex, of de Sá, Grant, Camargo, Heyer, Ponssa, and Stanley, 2014, S. Am. J. Herpetol., 9(Spec. Issue 1): 1–123, who provided a summary of the relevant literature of this species (adult and larval morphology, identification, advertisement call, and range) on p. 29–30, and who implied that previous records from Peru are assigned to other species. Schulze, Jansen, and Köhler, 2015, Zootaxa, 4016: 72–74, described, diagnosed, and pictured the larva. Vicente-Ferreira, Nascimento, Batista, Kardush, Reyes, and Garey, 2024, Biota Neotrop., 24(1: e20231526): 1–17, provided records from the Refúgio Biológico Bela Vista, Paraná, southern Brazil (adjacent to the Paraguay border), as well as providing identification keys to these species based on larval and adult morphology.

External links:

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For access to general information see Wikipedia
For additional sources of general information from other websites search Google
For access to relevant technical literature search Google Scholar
For images search CalPhoto Images and Google Images
To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.