Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Leptodactylus intermedius Lutz, 1930

Class: Amphibia > Order: Anura > Family: Leptodactylidae > Subfamily: Leptodactylinae > Genus: Leptodactylus > Species: Leptodactylus intermedius

Leptodactylus intermedius Lutz, 1930, Mem. Oswaldo Cruz, Rio de Janeiro, 23: 8, 27. Syntypes: AL-MNRJ 1438-1441, according to Heyer, 1970, Contrib. Sci. Nat. Hist. Mus. Los Angeles Co., 191: 18. Type locality: "Manacapui [= Manacapurú] perto do Manaos", Amazonas, Brazil (Portuguese text); "Manacapuri near Manaos" Brazil (English text).

English Names

None noted.

Distribution

Lowlands of the Amazon River Basin in Brazilian Amazonia (including at least parts of Rondônia, Amazonas, Amapá, and Pará), occurring also in Bolivia (Pando and Santa Cruz) and in the Eastern Guiana Shield in eastern French Guiana.

Geographic Occurrence

Natural Resident: Bolivia, Brazil, French Guiana

Comment

Removed from the synonymy of Leptodactylus petersii by Gazoni, Lyra, Ron, Strüssmann, Baldo, Narimatsu, Pansonato, Schneider, Giaretta, Haddad, Parise-Maltempi, and Carvalho, 2021, Zool. Anz., 290: 127 (who discussed morphology, molecular markers, vocalizations, and cytogenetics), where it had been placed by Heyer, 1994, Smithson. Contrib. Zool., 546: 96 (and previously in the synonymy of Leptodactylus wagneri according to Heyer, 1970, Contrib. Sci. Nat. Hist. Mus. Los Angeles Co., 191: 18). De la Riva, Köhler, Lötters, and Reichle, 2000, Rev. Esp. Herpetol., 14: 42, provided the Bolivian record (as Leptodactylus petersii). Carvalho, Fouquet, Lyra, Giaretta, Costa-Campos, Rodrigues, Haddad, and Ron, 2022, Syst. Biodiversity, 20 (1: 2089269): 1–31, reported on the systematics, phylogenetics, and advertisement call. Taucce, Costa-Campos, Carvalho, and Michalski, 2022, Eur. J. Taxon., 836: 96–130, reported on distribution, literature, and conservation status for Amapá, Brazil.

External links:

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For information on distribution, habitat, and conservation see the Map of Life
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For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.