Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Gephyromantis plicifer (Boulenger, 1882)

Class: Amphibia > Order: Anura > Family: Mantellidae > Subfamily: Mantellinae > Genus: Gephyromantis > Species: Gephyromantis plicifer

Rana plicifera Boulenger, 1882, Cat. Batr. Sal. Coll. Brit. Mus., Ed. 2: 464. Syntypes: BMNH (4 specimens), by original designation; BMNH 82.3.16.56-59 according to Blommers-Schlösser and Blanc, 1991, Faune de Madagascar, 75: 210. Type locality: "East Betsileo", Madagascar.

Mantidactylus pliciferus — Boulenger, 1895, Ann. Mag. Nat. Hist., Ser. 6, 15: 450; Mocquard, 1895, Bull. Soc. Philomath., Paris, Ser. 8, 7: 129. Incorrect subsequent spelling.

Mantidactylus sculpturatus Ahl, 1929 "1928", Mitt. Zool. Mus. Berlin, 14: 477. Holotype: ZMB unnumbered according to the original publication; ZMB 30515 according to Vences and Glaw, 2001, Alytes, 19: 129; reported lost, apparently in error, by Blommers-Schlösser and Blanc, 1991, Faune de Madagascar, 75: 210. Type locality: "Nord-West-Madagascar"; this locality considered likely erroneus by Vences and Glaw, 2001, Alytes, 19: 129. Synonymy by Vences, Köhler, Andreone, Craul, Crottini, Du Preez, Preick, Rancilhac, Rödel, Scherz, Streicher, Hofreiter, and Glaw, 2021, Spixiana, München, 44: 175.

Mantidactylus (Mantidactylus) sculpturatus — Ahl, 1931, Das Tierreich, 55: 30.

Mantidactylus (Mantidactylus) pliciferus — Ahl, 1931, Das Tierreich, 55: 30. Incorrect subsequent spelling.

Mantidactylus (Gephyromantis) sculpturatus — Vences and Glaw, 2001, Alytes, 19: 129, by implication.

Gephyromantis sculpturatus — Vences and Glaw, 2006, in Vences et al. (eds.), Calls Frogs Madagascar: 20.

Gephyromantis (Duboimantis) sculpturatus — Glaw and Vences, 2006, Organisms Divers. Evol., 6: 244, by implication; Glaw and Vences, 2006, Organisms Divers. Evol., Electron. Suppl., 11(1): 2.

Mantidactylus (Gephyromantis) plicifer — Dubois, 1992, Bull. Mens. Soc. Linn. Lyon, 61: 312.

Gephyromantis plicifer — Vences and Glaw, 2006, in Vences et al. (eds.), Calls Frogs Madagascar: 20.

Gephyromantis (Duboimantis) plicifer — Glaw and Vences, 2006, Organisms Divers. Evol., 6: 244, by implication; Glaw and Vences, 2006, Organisms Divers. Evol., Electron. Suppl., 11(1): 2.

Common Names

Common Madagascar Frog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 101).

Distribution

East-central Madagascar.

Geographic Occurrence

Natural Resident: Madagascar

Endemic: Madagascar

Comment

Prior to the revision of Glaw and Vences, 2006, Organisms Divers. Evol., 6: 236-253, in the Mantidactylus (Gephyromantis) granulatus group of Dubois, 1992, Bull. Mens. Soc. Linn. Lyon, 61: 312. Subsequently, Vences and Glaw, 2001, Alytes, 19: 127-129, provided an account and placed this species in their Mantidactylus asper group. Glaw and Vences, 2007, Field Guide Amph. Rept. Madagascar, Ed. 3: 222-223, provided an account. See statement of geographic range, habitat, and conservation status (as Mantidactylus plicifer) in Stuart, Hoffmann, Chanson, Cox, Berridge, Ramani, and Young, 2008, Threatened Amph. World: 624. Randrianiaina, Wollenberg, Hiobiarilanto, Strauss, Glos, and Vences, 2011, J. Nat. Hist., London, 45: 2871–2900, described larval morphology (as Gephyromantis sculpturatus). Rodriguez, Börner, Pabijan, Gehara, Haddad, and Vences, 2015, Evol. Ecol., 29: 765–785 (and supplemental data), reported (as Gephyromantis sculpturatus) on phylogeographic diversity and its causes. Vences, Köhler, Andreone, Craul, Crottini, Du Preez, Preick, Rancilhac, Rödel, Scherz, Streicher, Hofreiter, and Glaw, 2021, Spixiana, München, 44: 175–202, provided a molecular analysis that showed that the name Gephyromantis plicifer actually not to populations formerly associated with the name, but with a candidate species formerly referred to as Gephyromantis sp. Ca21, and that Gephyromantis sculpturatus is a synonym. Their data also required that Gephyromantis pedronoi be recognized for populations in southeastern Madagascar formerly associated with this species.

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For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.