Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Leptobrachella gracilis (Günther, 1872)

Class: Amphibia > Order: Anura > Family: Megophryidae > Subfamily: Leptobrachiinae > Genus: Leptobrachella > Species: Leptobrachella gracilis

Leptobrachium gracile Günther, 1872, Proc. Zool. Soc. London, 1872: 598. Holotype: BMNH 1872.2.19.35 according to Dehling, 2012, Zootaxa, 3328: 22; reregiostered as BMnh 1947.2.19.35 acccording to museum records. Type locality: "Matang", Sarawak, Malaysia (Borneo).

Leptobrachium gracile — Boulenger, 1882, Cat. Batr. Sal. Coll. Brit. Mus., Ed. 2: 440.

Megalophrys gracilis — Boulenger, 1908, Proc. Zool. Soc. London, 1908: 421.

Megophrys gracilis — Smith, 1930, Bull. Raffles Mus., 3: 133.

Leptobrachium gracilis — Berry and Hendrickson, 1963, Copeia, 1963: 643; Inger, 1966, Fieldiana, Zool., 52: 24.

Leptobrachium (Leptolalax) gracile — Dubois, 1980, Bull. Mens. Soc. Linn. Lyon, 49: 476.

Leptobrachium (Leptolalax) gracilis — Inger, 1983, in Rhodin and Miyata (eds.), Adv. Herpetol. Evol. Biol.: 16.

Leptolalax gracilis — Dubois, 1983, Alytes, 2: 148.

Leptolalax (Leptolalax) gracilis — Delorme, Dubois, Grosjean, and Ohler, 2006, Alytes, 24: 15.

Leptobrachella gracilis — Chen, Poyarkov, Suwannapoom, Lathrop, Wu, Zhou, Yuan, Jin, Chen, Liu, Nguyen, Nguyen, Duong, Eto, Nishikawa, Matsui, Orlov, Stuart, Brown, Rowley, Murphy, Wang, and Che, 2018, Mol. Phylogenet. Evol., 124: 162, by implication.

English Names

Matang Asian Toad (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 86).

Gracile Litter Frog (Chan-ard, 2003, Photograph. Guide Amph. Thailand: 78).

Sarawak Slender Litter Frog (Das, Jankowski, Makmor, and Haas, 2007, Mitt. Hamburg. Zool. Mus. Inst., 104: 154).

Slender Litter Frog (Dehling, 2012, Zootaxa, 3328: 21).

Distribution

Northern Borneo in Kalimantan (Indonesia), Brunei, and Sarawak (Malaysia); see comment.

Geographic Occurrence

Natural Resident: Brunei, Indonesia, Malaysia, Malaysia, East (Sarawak and/or Sabah)

Comment

See accounts by Bourret, 1942, Batr. Indochine: 206-207; Inger, 1966, Fieldiana, Zool., 52: 24-28; Berry, 1975, Amph. Fauna Peninsular Malaysia: 32-33, and Inger, Stuebing, and Tan, 1995, Raffles Bull. Zool., 43: 121-122. Matsui, 1997, Copeia, 1997: 158–165, reported on advertisement call. See identification table by Manthey and Grossmann, 1997, Amph. Rept. Südostasiens: 68-69, to compare this species with other megophryids in the Sunda Shelf region. Das, Jankowski, Makmor, and Haas, 2007, Mitt. Hamburg. Zool. Mus. Inst., 104: 154, provided a brief description. See statement of geographic range, habitat, and conservation status in Stuart, Hoffmann, Chanson, Cox, Berridge, Ramani, and Young, 2008, Threatened Amph. World: 625. Das, 2007, Amph. Rept. Brunei: 36, provided a photograph and a brief account. Dehling, 2012, Zootaxa, 3328: 20-34, reviewed and rediagnosed the systematics of the species, and concluded that Leptolalax gracilis proper is restricted to Borneo and that Malay Peninsula populations represent two unnamed species (on now named Leptobrachella platycephala) conflated with Leptobrachella gracilis. Matsui, 2006, Zool. Sci., Tokyo, 23: 821-830, noted that the isolated record in western Thailand (Thong Pha Phum Forest, Kanchanaburi Province) by Chan-ard, 2003, Photograph. Guide Amph. Thailand: 78-79, is likely referable to Leptobrachella melanoleuca. The Gunung Benom population is now recognized as Leptobrachella platycephala. The Gunung Tahan population previously referred to this species remains unnamed.

External links:

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For additional sources of general information from other websites search Google
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For images search CalPhoto Images and Google Images
To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observation see iNaturalist
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.