Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Leptobrachella heteropus (Boulenger, 1900)

Class: Amphibia > Order: Anura > Family: Megophryidae > Subfamily: Leptobrachiinae > Genus: Leptobrachella > Species: Leptobrachella heteropus

Leptobrachium heteropus Boulenger, 1900, Ann. Mag. Nat. Hist., Ser. 7, 6: 186. Holotype: Selangor Mus. (now MNM), by original designation; BMNH records have 1947.2.25.7 (originally 1901.3.20.6) as the holotype, presumably on exchange from MNM. Type locality: "Larut Hills at an altitude of 3500 feet", Perak, Malaya, Malaysia.

Megalophrys heteropus — Boulenger, 1908, Proc. Zool. Soc. London, 1908: 422.

Megophrys heteropus — Smith, 1930, Bull. Raffles Mus., 3: 133.

Leptobrachium heteropus — Berry and Hendrickson, 1963, Copeia, 1963: 643.

Leptobrachium (Leptolalax) heteropus — Dubois, 1980, Bull. Mens. Soc. Linn. Lyon, 49: 476.

Leptolalax heteropus — Dubois, 1983, Alytes, 2: 148.

Leptolalax (Leptolalax) heteropus — Delorme, Dubois, Grosjean, and Ohler, 2006, Alytes, 24: 15.

Leptobrachella heteropus — Chen, Poyarkov, Suwannapoom, Lathrop, Wu, Zhou, Yuan, Jin, Chen, Liu, Nguyen, Nguyen, Duong, Eto, Nishikawa, Matsui, Orlov, Stuart, Brown, Rowley, Murphy, Wang, and Che, 2018, Mol. Phylogenet. Evol., 124: 162, by implication.

Common Names

Malaysian Asian Toad (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 86).

Variable Litter Frog (Chan-ard, 2003, Photograph. Guide Amph. Thailand: 80).

Malaysian Litter Toad (Poyarkov, Nguyen, Popov, Geissler, Pawangkhanant, Neang, Suwannapoom, and Orlov, 2021, Russ. J. Herpetol., 28 (3A): 30).

Distribution

Malay Peninsula, Malaysia, and adjacent southernmost peninsular Thailand (see comment).

Geographic Occurrence

Natural Resident: Malaysia, Malaysia, West (Peninsular), Thailand

Comment

See accounts by Bourret, 1942, Batr. Indochine: 207–208 (as Megophrys heteropus), and Berry, 1975, Amph. Fauna Peninsular Malaysia: 34. See brief account and photo by Manthey and Grossmann, 1997, Amph. Rept. Südostasiens: 76–77. Matsui, 1997, Copeia, 1997: 158–165, reported on advertisement call. Chan-ard, 2003, Photograph. Guide Amph. Thailand: 80–81, provided a very brief account, map for Thailand, and photograph. Chan-ard and Thong-aree, 2003, BRT Research Report, 2003: 245–258, reported the species from Hala-Bala Wildlife Sanctuary, Yala and Narathiwat province, Thailand. Matsui, 2006, Zool. Sci., Tokyo, 23: 821–830, suggested that populations of Leptolalax heteropus from Thailand may be referable to Leptolalax solus. Grismer, Chan, Grismer, Wood, and Ahmad, 2010, Russ. J. Herpetol., 17: 147–160, reported localities from the Banjaran Bintang Mountains, northwestern peninsular Malaysia. Niyomwan, Srisom, and Pawangkhanant, 2019, Field Guide Amph. Thailand: 122–123, provided a brief account (photograph, habitat, and range) for Thailand (in Thai).

External links:

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For access to general information see Wikipedia
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For images search CalPhoto Images and Google Images
To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.