Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Scutiger khumbu Hofmann, Ohler, Baniya, Dubois, Flecks, Jablonski, Schmidt, and Dufresnes, 2025

Class: Amphibia > Order: Anura > Family: Megophryidae > Subfamily: Leptobrachiinae > Genus: Scutiger > Species: Scutiger khumbu

Scutiger khumbu Hofmann, Ohler, Baniya, Dubois, Flecks, Jablonski, Schmidt, and Dufresnes, 2025, Eur. J. Taxon., 989: 203. Holotype: ZFMK 104174, by original designation. Type locality: "NEPAL • ♂; Bagmati Province, Dolakha District, Jiri; 27°41′48″ N, 86°16′30″ E (WGS 84); 3396 m a.s.l." ZooBank Publication registration: 8C2D01C4-2395-407F-9358-658ACDEBB853

Schutiger khumbu makalu Hofmann, Ohler, Baniya, Dubois, Flecks, Jablonski, Schmidt, and Dufresnes, 2025, Eur. J. Taxon., 989: 209. Holotype: MNHN 2003.3041, by original designation. Type locality: "NEPAL • ♂; Bhojpur District, Salpa Pokhari; 27°26′47″ N, 86°56′01″ E (WGS 84)". ZooBank Publication registration: 8C2D01C4-2395-407F-9358-658ACDEBB853

Common Names

None noted.

Distribution

Occurs in mountain forests in high-montane areas of the drainage area of the Koshi basin, northeastern Nepal, more specifically, along the Khumbu Himal and adjacent Makalu region between ca 2800 and 3900 m elevation; known from the Sagarmatha and Koshi Zone, specifically the Dolakha District (Jiri), Ramechhap District, Solu Khumbu District (Lamjura La, Paiya, Surkie La, Taksindu), Bhojpur District (Salpa Pokhari), and in the Sankhuwasabha District.

Geographic Occurrence

Natural Resident: Nepal

Endemic: Nepal

Comment

Comparative adult and larval morphology and molecular markers detailed in the original publication, where two subspecies were delimited, a dot map provided, and it was noted that this species had been previously confused with Scutiger sikimmensis.

External links:

Please note: these links will take you to external websites not affiliated with the American Museum of Natural History. We are not responsible for their content.

For access to general information see Wikipedia
For additional sources of general information from other websites search Google
For access to relevant technical literature search Google Scholar
For images search CalPhoto Images and Google Images
To search the NIH genetic sequence database, see GenBank
For related information on conservation and images as well as observations see iNaturalist