Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Dyscophinae Boulenger, 1882

Class: Amphibia > Order: Anura > Family: Microhylidae > Subfamily: Dyscophinae

3 species

Dyscophidae Boulenger, 1882, Cat. Batr. Sal. Coll. Brit. Mus., Ed. 2: 179. Type genus: Dyscophus Grandidier, 1872.

Dyscophinae — Gadow, 1901, Amphibia and Reptiles: 139.

Dyscophiinae — Kuhn, 1965, Die Amphib.: 84.

Dyscophidae — Bossuyt and Roelants, 2009, in Hedges and Kumar (eds.), Timetree of Life: 358.

Dyscophini — Dubois, Ohler, and Pyron, 2021, Megataxa, 5: 218. Tribe.

Common Names

None noted.

Distribution

Madagascar.

Comment

See comment under Asterophryinae. Parker, 1934, Monogr. Frogs Fam. Microhylidae, considered Dyscophinae (sensu lato, as including Calluella, now in Microhylinae) to be the stem-group from which the rest of Microhylidae (excluding Scaphiophryninae; by implication) was derived. The literature of this controversy was reviewed by Blommers-Schlösser, 1975, Beaufortia, 24: 7-26, who retained the African and Indo-Malayan genera together in this group. See Blommers-Schlösser, 1976, Genetica, 46: 199-210, for karyotypes. Frost, Grant, Faivovich, Bain, Haas, Haddad, de Sá, Channing, Wilkinson, Donnellan, Raxworthy, Campbell, Blotto, Moler, Drewes, Nussbaum, Lynch, Green, and Wheeler, 2006, Bull. Am. Mus. Nat. Hist., 297: 226, restricted Dyscophinae to Dyscophus and transferred Calluella to Microhylinae. Bossuyt and Roelants, 2009, in Hedges and Kumar (eds.), Timetree of Life: 358, considered this taxon a distinct family based on its ancient origin. Pyron and Wiens, 2011, Mol. Phylogenet. Evol., 61: 543-583, in their molecular study, suggested that Dyscophinae is the sister of Microhylinae.

Contained taxa (3 sp.):

Dyscophus Grandidier, 1872 (3 sp.)

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