Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Micryletta nigromaculata Poyarkov, Nguyen, Duong, Gorin, and Yang, 2018

Class: Amphibia > Order: Anura > Family: Microhylidae > Subfamily: Microhylinae > Genus: Micryletta > Species: Micryletta nigromaculata

Micryletta nigromaculata Poyarkov, Nguyen, Duong, Gorin, and Yang, 2018, PeerJ, 6 (e5771): 9. Holotype: ZMMU A5934, by original designation. Type locality: "from the limestone evergreen forest (20.8123N,106.9988E, at an elevation of 90 m a.s.l.), Cat Ba National Park, Hai Phong Province, northern Vietnam." Zoobank publication registration: 32150A60-5D04-4116-9816-0ED3E457504A.

Common Names

Black-spotted Paddy Frog (original publication).

Distribution

Known from limestone karst areas covered by primary tropical forest on either sides of the Red River Valley in northern Vietnam, specifically in Cat Ba National Park in Hai Phong Province, Cuc Phuong National Park in Ninh Binh Province, and Son La City and Phong Lai Commune in Son La Province, at 90–680 m elevation; see comment regarding Laos (Northern Annamites in Nakai District, Khammouan Province, 220 m) population.

Geographic Occurrence

Natural Resident: Vietnam

Likely/Controversially Present: Laos

Endemic: Vietnam

Comment

The sister taxon of the remaining species of Micryletta according to the original publication. Das, Garg, Hamidy, Smith, and Biju, 2019, PeerJ, 7(e7012): 1–32, suggested that their Micryletta cf. nigromaculata from Bolikhamxay, Thaphabat District, Bolikhamsai Province, northwest-central Laos represents a distinct species from those populations in Vietnam. Pham, Nguyen, Sung, Tran, Le, and Ngo, 2023, Acta Herpetol., Firenze, 18: 131–138, reported populations from Son La Province, northwestern Vietnam, and discussed habitat and dietary ecology. Stuart, Seateun, Sivongxay, and Phimmachak, 2025 "2024", in Wildlife Lao PDR: 37, summarized (as Micryletta cf. nigromaculata) literature, taxonomy, habitat, and detailed range for Laos.

External links:

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For access to general information see Wikipedia
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To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist