Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Nanohyla marmorata (Bain and Nguyen, 2004)

Class: Amphibia > Order: Anura > Family: Microhylidae > Subfamily: Microhylinae > Genus: Nanohyla > Species: Nanohyla marmorata

Microhyla marmorata Bain and Nguyen, 2004, Copeia, 2004: 508. Holotype: AMNH 161364, by original designation. Type locality: "Po-mu Mountain, Huong Son Reserve (now known as Huong Son Forestry and Service Company), Huong Son District, Rao An Region, Ha Tinh Province, Vietnam (18° 20′ 53″ N, 105° 14′ 38″ E, elevation 1010-1080 m)".

Microhyla pulverata Bain and Nguyen, 2004, Copeia, 2004: 513. Holotype: FMNH 254122, by original designation. Type locality: "Buon Luoi village, 20 km northwest of Kannack (Kannack coordinates: 14° 20′ N, 108° 36′ E), An Khe District, Gia Lai Province, Vietnam, (700-750 m elevation)". Synonymy by Gorin, Solovyeva, Hasan, Okamiya, Karunarathna, Pawangkhanant, de Silva, Juthong, Milto, Nguyen, Suwannapoom, Haas, Bickford, Das, and Poyarkov, 2020, PeerJ, 8 (e9411): 31.

Nanohyla marmorata — Gorin, Scherz, Korost, and Poyarkov, 2021, Zoosyst. Evol., 97: 38.

English Names

Marble Pigmy Frog (Nguyen, Ho, and Nguyen, 2005, Checklist Amph. Rept. Vietnam: 45).

Dusty Speckling Venter Pigmy Frog (Microhyla pulverata [no longer recognized]: Nguyen, Ho, and Nguyen, 2005, Checklist Amph. Rept. Vietnam: 46).

Marbled Pigmy Narrow-Mouth Frog (Poyarkov, Nguyen, Popov, Geissler, Pawangkhanant, Neang, Suwannapoom, and Orlov, 2021, Russ. J. Herpetol., 28 (3A): 43).

Distribution

Ha Tinh, Quang Binh, Quang Tri, Quang Nam, Kon Tum, Thanh Hoa, Gia Lai, Binh Dinh, and Thua Thien-Hue Provinces in Vietnam; also in adjacent Laos in Khammouane, Xekong, and provinces.

Comment

Confused with Microhyla (now Nanohyla) annamensis prior to its description according to the original publication. Nguyen, Ho, and Nguyen, 2005, Checklist Amph. Rept. Vietnam: 45, provided specific localities for Vietnam. Bain, Nguyen, and Doan, 2007, Herpetol. Rev., 38: 107, provided a record from Thua Thien-Hue Province, Vietnam, and briefly discussed the range. Pham, Nguyen, Hoang, and Ziegler, 2016, Herpetol. Notes, 9: 35, provided a record for Thanh Hoa Province, Vietnam, and commented on the range. Gorin, Solovyeva, Hasan, Okamiya, Karunarathna, Pawangkhanant, de Silva, Juthong, Milto, Nguyen, Suwannapoom, Haas, Bickford, Das, and Poyarkov, 2020, PeerJ, 8 (e9411): 1–47, placed this species in their Microhyla (now Nanohyla) annectens group. Brakels, Maury, Phimmachak, and Nguyen, 2021, Herpetol. Rev., 52: 792, provided records from Xaychamphone District, Bolikhamsai Province, and Dakcheung District, Xekong Province, southeastern Laos. Duong and Ngo, 2022, Herpetol. Notes, 15: 751–762, discussed range in Vietnam and provided records for Binh Dinh Province.

External links:

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For additional sources of information from other sites search Google
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To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For related information on conservation and images as well as observation see iNaturalist
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.