Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Nanohyla petrigena (Inger and Frogner, 1979)

Class: Amphibia > Order: Anura > Family: Microhylidae > Subfamily: Microhylinae > Genus: Nanohyla > Species: Nanohyla petrigena

Microhyla petrigena Inger and Frogner, 1979, Sarawak Mus. J., 27: 315. Holotype: FMNH 207705, by original designation. Type locality: "Nanga Tekalit, Kapit District, Sarawak", Malaysia (Borneo).

Microhyla (Microhyla) petrigena — Dubois, 1987, Alytes, 6: 3.

Nanohyla petrigena — Gorin, Scherz, Korost, and Poyarkov, 2021, Zoosyst. Evol., 97: 38.

English Names

Kapit Rice Frog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 91).

Pothole Narrow-mouthed Frog (Das, 2007, Amph. Rept. Brunei: 44).

Distribution

Northern Borneo in southeastern Sabah and central Sarawak (Malaysia), Brunei, and central Kalimantan (Indonesia); Sulu Archipelago of Philippines.

Comment

See Leong and Heok, 2000, Herpetol. Rev., 31: 109, for Indonesian record. See identification table (as Microhyla) by Manthey and Grossmann, 1997, Amph. Rept. Südostasiens: 46, to compare this species with other microhylids in the Sunda Shelf region. Malkmus, Manthey, Vogel, Hoffmann, and Kosuch, 2002, Amph. Rept. Mount Kinabalu: 132, provided an account. See statement of geographic range, habitat, and conservation status in Stuart, Hoffmann, Chanson, Cox, Berridge, Ramani, and Young, 2008, Threatened Amph. World: 626. See statement of geographic range, habitat, and conservation status in Stuart, Hoffmann, Chanson, Cox, Berridge, Ramani, and Young, 2008, Threatened Amph. World: 626. Das, 2007, Amph. Rept. Brunei: 44, provided a photograph and a brief account. Dehling, 2010, Salamandra, 46: 114-116, reported on the advertisement call. Haas, Kueh, Joseph, bin Asri, Das, Hagmann, Schwander, and Hertwig, 2018, Evol. Syst., 2: 89–114, provided a brief account of morphology and natural history for the Sabah population. Firdaus, Ratih, Karima, Kusuma, and Suastika, 2018, Bioinform. Biomed. Res. J., 1: 1–6, reported on the mtDNA phylogenetic relationships of the species of Microhyla within Sumatra, Java, Borneo, and peninsular Malaysia, placing this species as the sister of Microhyla perparva. Gorin, Solovyeva, Hasan, Okamiya, Karunarathna, Pawangkhanant, de Silva, Juthong, Milto, Nguyen, Suwannapoom, Haas, Bickford, Das, and Poyarkov, 2020, PeerJ, 8 (e9411): 1–47, placed this species in their (now Nanohyla) group.

External links:

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For additional sources of information from other sites search Google
For images search CalPhoto Images and Google Images
To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For related information on conservation and images as well as observation see iNaturalist; for a quick link to their maps see iNaturalist KML
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.