Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Heleioporus Gray, 1841

Class: Amphibia > Order: Anura > Superfamily: Myobatrachoidea > Family: Limnodynastidae > Genus: Heleioporus

6 species

Heleioporus Gray, 1841, Ann. Mag. Nat. Hist., Ser. 1, 7: 91. Type species: Heleioporus albopunctatus Gray, 1841, by monotypy.

Helioporus — Gray, 1841, in Grey (ed.), J. Two Exped. Discovery NW and W. Aust., 2(App. E): 447. Incorrect subsequent spelling.

Perialia Gray, 1845, in Eyre, J. Exped. Discov. Cent. Aust. 1840–1841, Append.: 406. Type species: Perialia eyrei Gray, 1845, by subsequent designation of Parker, 1940, Novit. Zool., 42: 1-106. Synonymy by Boulenger, 1882, Cat. Batr. Sal. Coll. Brit. Mus., Ed. 2: 271. Resurrected; without discussion, by Wells and Wellington, 1985, Aust. J. Herpetol., Suppl. Ser., 1: 3. See Tyler, 1985, Herpetol. Rev., 16: 69; Australian Society of Herpetologists, 1987, Bull. Zool. Nomencl., 44: 116-121; Tyler, 1988, Bull. Zool. Nomencl., 45: 152; Holthius, 1988, Bull. Zool. Nomencl., 45 and decision by ICZN (Anonymous, 1991, Bull. Zool. Nomencl., 48: 337-338).

Heleioforus — Krefft, 1865, Pap. Proc. R. Soc. Tasmania, 1865: 17. Incorrect subsequent spelling.

Philocryphus Fletcher, 1894, Proc. Linn. Soc. New South Wales, Ser. 2, 8: 233. Type species: Philocryphus flavoguttatus Fletcher, 1894, by monotypy. Synonymy by Nieden, 1923, Das Tierreich, 46: 525. Resurrected; without discussion, by Wells and Wellington, 1985, Aust. J. Herpetol., Suppl. Ser., 1: 3. See Tyler, 1985, Herpetol. Rev., 16: 69; Australian Society of Herpetologists, 1987, Bull. Zool. Nomencl., 44: 116-121; Tyler, 1988, Bull. Zool. Nomencl., 45: 152; Holthius, 1988, Bull. Zool. Nomencl., 45 and decision by ICZN (Anonymous, 1991, Bull. Zool. Nomencl., 48: 337-338).

Common Names

Giant Burrowing Frogs (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 94).

Distribution

Southern Western Australia, southwestern New South Wales, and eastern Victoria, Australia.

Comment

Reviewed by Lee, 1967, Aust. J. Zool., 15: 367-439. See Barker, Grigg, and Tyler, 1995, Field Guide Aust. Frogs., Ed. 2, for accounts and keys to species. See comment under Neobatrachus. Lynch, 1971, Misc. Publ. Mus. Nat. Hist. Univ. Kansas, 53: 79, suggested the relationship Heleioporus + (Neobatrachus + Notaden). Cogger, 2018, Rept. Amph. Australia, 7th ed.: 40–43, provided accounts, photographs, and polygon distribution maps.

Contained taxa (6 sp.):

Heleioporus albopunctatus Gray, 1841

Heleioporus australiacus (Shaw and Nodder, 1795)

Heleioporus barycragus Lee, 1967

Heleioporus eyrei (Gray, 1845)

Heleioporus inornatus Lee and Main, 1954

Heleioporus psammophilus Lee and Main, 1954

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