Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Phrynobatrachus batesii (Boulenger, 1906)

Class: Amphibia > Order: Anura > Family: Phrynobatrachidae > Genus: Phrynobatrachus > Species: Phrynobatrachus batesii

Arthroleptis batesii Boulenger, 1906, Ann. Mag. Nat. Hist., Ser. 7, 17: 318. Syntypes: BMNH 1947.2.6.76–77 (formerly 1902.11.12.86-87) and 1947.2.6.78-80 (formerly 1906.5.28.105-107), according to Grandison, 1985, in Frost (ed.), Amph. Species World: 444, and museum records. Type locality: "Efulen and .... Zima [= Sangmelima]", Cameroon.

Pseudarthroleptis batesi — Deckert, 1938, Sitzungsber. Ges. Naturforsch. Freunde Berlin, 1938: 159.

Phrynobatrachus batesi — Perret, 1959, Bull. Soc. Neuchatel. Sci. Nat., 82: 251.

Common Names

Bates' River Frog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 103).

Bates' Puddle Frog (Channing and Rödel, 2019, Field Guide Frogs & Other Amph. Afr.: 284).

Distribution

Southeastern Nigeria and western Cameroon through Equatorial Guinea to Gabon (see comment); isolated and questionable record from Ghana (see comment) and southwestern Nigeria.

Geographic Occurrence

Natural Resident: Cameroon, Equatorial Guinea, Gabon, Nigeria

Likely/Controversially Present: Ghana

Comment

See account in Perret, 1966, Zool. Jahrb., Jena, Abt. Syst., 93: 359–360. The Ghana records by Schiøtz, 1964, Vidensk. Medd. Dansk Naturhist. Foren., 127: 1–17, may be based on another species, according to J.-L. Perret (personal communication to DRF). A form close to this species was reported from Gabon by Frétey and Blanc, 2002 "2001", Bull. Soc. Zool. France, 126: 386. Schiøtz, 1963, Vidensk. Medd. Dansk Naturhist. Foren., 125: 41, provided records for Nigeria. Frétey, 2008, Alytes, 25: 99–172, summarized the literature. Rödel, Ofori-Boateng, Penner, and Hillers, 2009, Zootaxa, 1970: 58, regarded the Ghana records of this species to be based on a misidentification. In species group C of Zimkus, Rödel, and Hillers, 2010, Mol. Phylogenet. Evol., 55: 883–900. Dewynter and Frétey, 2019, Cah. Fondation Biotope, 27: 31, summarized the literature for Gabon and suggested that although the species probably does get to Gabon, that all records previous assigned to this country are based on misidentifications. Channing and Rödel, 2019, Field Guide Frogs & Other Amph. Afr.: 284–285, provided a brief account, photograph, and range map (which excluded Ghana and Nigeria). In the Phrynobatrachus natalensis group of Goutte, Reyes-Velasco, and Boissinot, 2019, ZooKeys, 824: 53–70. In the Phrynobatracus werneri group according to Gvoždík, Nečas, Dolinay, Zimkus, Schmitz, and Fokam, 2020, PeerJ, 8 (e8393): 1–52. See brief account, range map for Equatorial Guinea in Sánchez-Vialas, Calvo-Revuelta, Castroviejo-Fisher, and De la Riva, 2020, Proc. California Acad. Sci., Ser. 4, 66: 137–230.

External links:

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For access to general information see Wikipedia
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For images search CalPhoto Images and Google Images
To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.