Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Phrynobatrachus calcaratus (Peters, 1863)

Class: Amphibia > Order: Anura > Family: Phrynobatrachidae > Genus: Phrynobatrachus > Species: Phrynobatrachus calcaratus

Hemimantis calcaratus Peters, 1863, Monatsber. Preuss. Akad. Wiss. Berlin, 1863: 451. Syntypes: ZMB 4923 and 52358, according to Bauer, Günther, and Klipfel, 1995, in Bauer et al. (eds.), Herpetol. Contr. W.C.H. Peters: 49. Type locality: "Boutry", Ghana. Placed on the Official List of Specific Names in Zoology by Opinion 1921, Anonymous, 1999, Bull. Zool. Nomencl., 56: 96-100.

Arthroleptis calcarata — Peters, 1875, Monatsber. Preuss. Akad. Wiss. Berlin, 1875: 210. .

Arthroleptis calcaratus — Boulenger, 1882, Cat. Batr. Sal. Coll. Brit. Mus., Ed. 2: 118.

Hyperolius (Arthroleptis) calcaratus — Knauer, 1878, Naturgesch. Lurche: 111.

Pseudarthroleptis calcaratus — Deckert, 1938, Sitzungsber. Ges. Naturforsch. Freunde Berlin, 1938: 158.

Phrynobatrachus (Pararthroleptis) calcaratus — Laurent, 1941, Rev. Zool. Bot. Afr., 34: 206.

Phrynobatrachus calcaratus — Guibé and Lamotte, 1958, Mem. Inst. Franç. Afr. Noire, 53: 257. Guibé and Lamotte, 1963, Mem. Inst. Franç. Afr. Noire, 66: 601-627

English Names

Boutry River Frog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 103).

Peters' Puddle Frog (Jacobsen, 2009, Afr. Herp News, 47: 5).

Distribution

Senegal, Guinea-Bissau, Guinea, and Liberia to Nigeria and through Cameroon to east-central Central African Republic and Bioko I. (Equatorial Guinea), below 1200 m elevation; expected in Gambia and Sierra Leone.

Geographic Occurrence

Natural Resident: Benin, Cameroon, Central African Republic, Equatorial Guinea, Ghana, Guinea, Guinea-Bissau, Ivory Coast, Liberia, Nigeria, Senegal, Togo

Likely/Controversially Present: Gambia, Sierra Leone

Comment

Reviewed by Guibé and Lamotte, 1963, Mem. Inst. Franç. Afr. Noire, 66: 601–627; and Perret, 1988, Arch. Sci., Genève, 41: 277–279. Rödel, 2000, Herpetofauna W. Afr., 1: 176–180, provided an account. Schiøtz, 1963, Vidensk. Medd. Dansk Naturhist. Foren., 125: 46–47, provided records for Nigeria. Emms, Jambang, Bah, Mankali, Rödel, and Barnett, 2005, Herpetol. Bull., London, 94: 6–16, provided a record for this or a very similar species from Gambia. Pauwels and Rödel, 2007, Herpetozoa, Wien, 19: 135–148, did not note this species as occurring in Gabon. Rödel, Brede, Schiefenhövel, Penner, Sinsin, and Nago, 2007, Salamandra, 43: 231–238, reported this species from the Lama Forest of southern Benin. Frétey, 2008, Alytes, 25: 99–172, summarized the literature. Hillers, Ofori-Boateng, Segniagbeto, Agyei, and Rödel, 2009, Zoosyst. Evol., Berlin, 85: 127–141, suggested that the name of this taxon covers a species complex. Jacobsen, 2009, Afr. Herp News, 47: 2–20, reported this species from east-north-central Central African Republic. In species group B of Zimkus, Rödel, and Hillers, 2010, Mol. Phylogenet. Evol., 55: 883–900, who noted this nominal species is a complex. Channing, Rödel, and Channing, 2012, Tadpoles of Africa: 275–276, provided information on comparative larval morphology. Auliya, Wagner, and Böhme, 2012, Bonn Zool. Bull., 61: 255–281, reported the species from the Bijagós Archipelago, Guinea-Bissau. Rödel and Glos, 2019, Zoosyst. Evol., 95: 24, reported this species from the Krahn-Bassa Proposed Protected Area in southeastern Liberia and commented on identification and habitat preference. Channing and Rödel, 2019, Field Guide Frogs & Other Amph. Afr.: 298–299, provided a brief account, photograph, and range map. See brief account and range map for Equatorial Guinea in Sánchez-Vialas, Calvo-Revuelta, Castroviejo-Fisher, and De la Riva, 2020, Proc. California Acad. Sci., Ser. 4, 66: 137–230. In the Phrynobatrachus calcaratus group of Goutte, Reyes-Velasco, and Boissinot, 2019, ZooKeys, 824: 53–70.

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For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.