Phrynobatrachus cornutus (Boulenger, 1906)

Class: Amphibia > Order: Anura > Family: Phrynobatrachidae > Genus: Phrynobatrachus > Species: Phrynobatrachus cornutus

Arthroleptis cornutus Boulenger, 1906, Ann. Mag. Nat. Hist., Ser. 7, 17: 318. Syntypes: BMNH 1947.2.30.49–50 (formerly 1906.5.28.103–104), according to Grandison, 1985, in Frost (ed.), Amph. Species World: 444, and museum records. Type locality: "Zima" (= Sangmelima, Départment Dja-et-Lobo), Cameroon. See comment by Frétey, Dubois, and Ohler, 2014, Alytes, 30: 44. 

Micrarthroleptis cornutusDeckert, 1938, Sitzungsber. Ges. Naturforsch. Freunde Berlin, 1938: 159.

Phrynobatrachus (Micrarthroleptis) cornutusLaurent, 1941, Rev. Zool. Bot. Afr., 34: 208.

Phrynobatrachus cornutusGuibé and Lamotte, 1958, Bull. Mus. Natl. Hist. Nat. Paris, Ser. 2, 30: 257; Guibé and Lamotte, 1963, Mem. Inst. Franç. Afr. Noire, 66: 626; Mertens, 1965, Bonn. Zool. Beitr., 16: 22.

Phrynobatrachus cornutus cornutusPerret, 1966, Zool. Jahrb., Jena, Abt. Syst., 93: 365.

English Names

Horned River Frog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 103).

Horned Puddle Frog (Jacobsen, 2009, Afr. Herp News, 47: 5).


Western Cameroon to northwestern Dem. Rep. Congo; Bioko and Equatorial Guinea through Gabon to north-east-central Central African Republic; likely into northwestern Angola.

Geographic Occurrence

Natural Resident: Cameroon, Central African Republic, Congo, Democratic Republic of the, Congo, Republic of the, Equatorial Guinea, Gabon

Likely/Controversially Present: Angola


Discussed by Perret, 1988, Arch. Sci., Genève, 41: 279–281. See Largen and Dowsett-Lemaire, 1991, Tauraco Res. Rept., 4: 151–152, for Rep. Congo records. Frétey and Blanc, 2002 "2001", Bull. Soc. Zool. France, 126: 386, reported this species from Gabon. Frétey, 2008, Alytes, 25: 99–172, summarized the literature. Jacobsen, 2009, Afr. Herp News, 47: 2–20, reported this species from east-north-central Central African Republic. In species group B of Zimkus, Rödel, and Hillers, 2010, Mol. Phylogenet. Evol., 55: 883–900. Nagy, Chifundera, Collet, and Gvoždík, 2013, Herpetol. Notes, 6:  413–419, provided a record from Bas-Congo, southwestern Dem. Rep. Congo. Baptista, Conradie, Vaz Pinto, and Branch, 2019, In Huntley, Russo, Lages, and Ferrand (eds.), Biodiversity in Angola: 256, noted specimens in Luki, Dem. Rep. Congo, near Angola (both the Cabinda enclave and northwestern Angola proper), suggesting its possible occurrence there. Dewynter and Frétey, 2019, Cah. Fondation Biotope, 27: 33, summarized the literature for Gabon and provided a photograph (p. 61). Channing and Rödel, 2019, Field Guide Frogs & Other Amph. Afr.: 298–299, provided a brief account, photograph, and range map. In the Phrynobatrachus calcaratus group of Goutte, Reyes-Velasco, and Boissinot, 2019, ZooKeys, 824: 53–70. See brief account and range map for Equatorial Guinea in Sánchez-Vialas, Calvo-Revuelta, Castroviejo-Fisher, and De la Riva, 2020, Proc. California Acad. Sci., Ser. 4, 66: 137–230. 

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