Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Phrynobatrachus inexpectatus Largen, 2001

Class: Amphibia > Order: Anura > Family: Phrynobatrachidae > Genus: Phrynobatrachus > Species: Phrynobatrachus inexpectatus

Phrynobatrachus inexpectatus Largen, 2001, Tropical Zool., 14: 301. Holotype: BMNH 1975.1711, by original designation. Type locality: "70 km NW of Kebre Mengist, near Bore (06° 23′ N 38° 35′ E, 2650 m), Ethiopia".

Common Names

Largen's Dwarf Puddle Frog (Largen and Spawls, 2010, Amph. Rept. Ethiopia Eritrea: 208; Spawls, Mazuch, and Mohammad, 2023, Handb. Amph. Rept. NE Afr.: 90).

Bore Puddle Frog (Channing and Rödel, 2019, Field Guide Frogs & Other Amph. Afr.: 310).

Distribution

Humid montane forests in southwestern Ethiopia west of the Rift Valley from near Dorse at 2600 m elevation) and east of the Rift Valley from near Bore (2650 m elevation) and from several nearby localities in the Harenna Forest of the Bale Mountains at 1400 to 2400 m elevation. See comment.

Geographic Occurrence

Natural Resident: Ethiopia

Endemic: Ethiopia

Comment

In the dwarf Phrynobatrachus group according to the original publication and compared with Phrynobatrachus scapularis, Phrynobatrachus albifer, Phrynobatrachus rouxi, Phrynobatrachus ukingensis, Phrynobatrachus mababiensis, Phrynobatrachus cryptotis. Frétey, 2008, Alytes, 25: 99-17, summarized the literature. In species group B of Zimkus, Rödel, and Hillers, 2010, Mol. Phylogenet. Evol., 55: 883-900. See account, photograph, and map for Ethiopia by Largen and Spawls, 2010, Amph. Rept. Ethiopia Eritrea: 208-209. Channing and Rödel, 2019, Field Guide Frogs & Other Amph. Afr.: 310–311, provided a brief account, photograph, and range map. In the Phrynobatrachus calcaratus group of Goutte, Reyes-Velasco, and Boissinot, 2019, ZooKeys, 824: 53–70. Tiutenko and Zinenko, 2020, Salamandra, 56: 135–147, commented on the identification with respect to Phrynobatrachus minutus and range (noting that the earlier record from Boka village, Kaffa province, Ethiopia, is based on juvenile Phrynobatrachus cf. natalensis), call, tadpole morphology, and breeding biology. Kassie Teme, Bekele Simegn, and Afework Bogale, 2022, Global Ecol. Conserv., 38 (e02211): 1–12, reported the species from Keffa Zone, southwestern Ethiopia, and discussed habitat. Spawls, Mazuch, and Mohammad, 2023, Handb. Amph. Rept. NE Afr.: 90, provided an account addressing identification, natural history, conservation status, and range, including a map.

External links:

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For access to general information see Wikipedia
For additional sources of general information from other websites search Google
For access to relevant technical literature search Google Scholar
For images search CalPhoto Images and Google Images
To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.