Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Phrynobatrachus leveleve Uyeda, Drewes, and Zimkus, 2007

Class: Amphibia > Order: Anura > Family: Phrynobatrachidae > Genus: Phrynobatrachus > Species: Phrynobatrachus leveleve

Phrynobatrachus leveleve Uyeda, Drewes, and Zimkus, 2007, Proc. California Acad. Sci., Ser. 4, 58: 371. Holotype: CAS 218901, by original designation. Type locality: "SãO TOMÉ AND PRÍNCIPE: São Tomé Island, Caxueira, along Agua Pete Pete, 0°18′N, 6°44′E, elevation 50 m."

Common Names

Calm Puddle Frog (Channing and Rödel, 2019, Field Guide Frogs & Other Amph. Afr.: 304).

Distribution

São Tomé and Rolas Islands, São Tomé and Principe, Gulf of Guinea.

Geographic Occurrence

Natural Resident: São Tomé and Principe

Endemic: São Tomé and Principe

Comment

Confused with Phrynobatrachus dispar prior to its naming according to the original publication. Frétey, 2008, Alytes, 25: 99-172, summarized the literature. In species group B of Zimkus, Rödel, and Hillers, 2010, Mol. Phylogenet. Evol., 55: 883-900. Channing and Rödel, 2019, Field Guide Frogs & Other Amph. Afr.: 304–305, provided a brief account, photograph, and range map. Bell, Ceríaco, Scheinberg, and Drewes, 2022, in Ceríaco et al. (eds.), Biodiversity Gulf Guinea Islands: 494–495, summarized the literature and systematics of the species. Ceríaco, Marques, Sousa, Veríssimo, Beja, and Ferreira, 2023, ZooKeys, 1168: 41–75, included this species in a report, key, and DNA barcode reference library for the amphibians of São Tomé and Príncipe. Sousa, Santos, Parrinha, Barbosa, Marques, Passos, and Ceríaco, 2026, Nat. Hist. Coll. Mus., 3(e184711): 1–63, discussed collecting effort on São Thome I., and nearby islands, and provided a grid map of this species.

External links:

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For access to general information see Wikipedia
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For images search CalPhoto Images and Google Images
To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For related information on conservation and images as well as observations see iNaturalist