Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Phrynobatrachus perpalmatus Boulenger, 1898

Class: Amphibia > Order: Anura > Family: Phrynobatrachidae > Genus: Phrynobatrachus > Species: Phrynobatrachus perpalmatus

Phrynobatrachus perpalmatus Boulenger, 1898, Proc. Zool. Soc. London, 1898: 479. Syntypes: BMNH 1947.2.4.34–35 (formerly 98.7.22.24–25), according to Grandison, 1985, in Frost (ed.), Amph. Species World: 449. Type locality: "about Lake Mwero", Zambia-Dem. Rep. Congo boundary.

Phrynobatrachus perpalmatus perpalmatus — Ahl, 1924, Mitt. Zool. Mus. Berlin, 11: 273.

Phrynobatrachus perpalmatus werneri Ahl, 1924, Mitt. Zool. Mus. Berlin, 11: 273. Syntypes: Presumably NHMW; by implication of referring to material reported by Werner, 1919, Denkschr. Akad. Wiss. Wien, Math. Naturwiss. Kl., 96: 19, although not noted in recent type lists . Type localities: "El Grassi und Weisser Nil (Tonga, Renk, Kashawal)". Secondary homonym of Arthroleptis werneri Nieden, 1910. Synonymy by Loveridge, 1933, Bull. Mus. Comp. Zool., 74: 346.

Phrynobatrachus (Phrynobatrachus) perpalmatus — Laurent, 1941, Rev. Zool. Bot. Afr., 34: 199.

English Names

Lake Mwero River Frog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 104).

Webbed Puddle Frog (Channing, 2001, Amph. Cent. S. Afr.: 310; Channing and Howell, 2006, Amph. E. Afr.: 288).

Distribution

South-central Sudan along the Nile River to central Dem. Rep. Congo, extreme western Tanzania, Zambia, Malawi, and northern Mozambique, possibly into northern Zimbabwe.

Geographic Occurrence

Natural Resident: Burundi, Congo, Democratic Republic of the, Malawi, Mozambique, South Sudan, Sudan, Tanzania, Zambia

Likely/Controversially Present: Zimbabwe

Comment

See accounts by Schmidt and Inger, 1959, Explor. Parc Natl. Upemba, Miss. G.F. de Witte, 56: 165–169, Inger, 1968, Explor. Parc Natl. Garamba, Miss. H. de Saeger, 52: 106–108, and Channing, 2001, Amph. Cent. S. Afr.: 310–311. Loveridge's records for Mozambique and Malawi are based on Phrynobatrachus acridoides; see Poynton, 1964, Ann. Natal Mus., 17: 140–141. See Poynton and Broadley, 1985, Ann. Natal Mus., 27: 158. Records of Phrynobatrachus perpalmatus from Cameroon by Perret and Mertens, 1957, Rev. Suisse Zool., 64: 73–78, are based on specimens of Phrynobatrachus steindachneri according to J.-L. Perret (personal communication). Channing and Howell, 2006, Amph. E. Afr.: 288–289, provided an account. Frétey, 2008, Alytes, 25: 99–172, summarized the literature and reported the species as occurring in Burundi, but without a specific locality. Not addressed by Zimkus, Rödel, and Hillers, 2010, Mol. Phylogenet. Evol., 55: 883–900, so not assigned to species group. Mercurio, 2011, Amph. Malawi: 231, provided an account for Malawi. Channing and Rödel, 2019, Field Guide Frogs & Other Amph. Afr.: 272–273, provided a brief account, photograph, and range map. Gvoždík, Nečas, Dolinay, Zimkus, Schmitz, and Fokam, 2020, PeerJ, 8 (e8393): 1–52, discussed misidentifications.

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For information on distribution, habitat, and conservation see the Map of Life
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For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.