Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Phrynobatrachus scheffleri (Nieden, 1911)

Class: Amphibia > Order: Anura > Family: Phrynobatrachidae > Genus: Phrynobatrachus > Species: Phrynobatrachus scheffleri

Arthroleptis scheffleri Nieden, 1911 "1910", Sitzungsber. Ges. Naturforsch. Freunde Berlin, 1910: 438. Syntypes: ZMB (169 specimens), by original designation. Type localities: "Kibwesi in Britisch-ostafrika", "Nairobi", "Zanzibar", and "Mpapua".

Arthroleptis albifer Ahl, 1924, Arch. Naturgesch., Abt. A,, 90: 251. Holotype: ZMB unnumbered, according to the original publication. Type locality: "Usaramo, Deutsch-Ost-Afrika", Tanzania. Synonymy with Phrynobatrachus minutus by Loveridge, 1929, Bull. U.S. Natl. Mus., 151: 107. This synonymy questioned by Largen, 2001, Tropical Zool., 14: 300. Synonymy (tentative) with Phrynobatrachus scheffleri by Schick, Zimkus, Channing, Köhler, and Lötters, 2010, Salamandra, 46: 29.

Pseudarthroleptis scheffleri — Deckert, 1938, Sitzungsber. Ges. Naturforsch. Freunde Berlin, 1938: 159.

Pseudarthroleptis albifer — Deckert, 1938, Sitzungsber. Ges. Naturforsch. Freunde Berlin, 1938: 158.

Phrynobatrachus (Pararthroleptis) scheffleri — Laurent, 1941, Rev. Zool. Bot. Afr., 34: 205.

Phrynobatrachus scheffleri — Largen, 2001, Tropical Zool., 14: 300.

Phrynobatrachus albifer — Largen, 2001, Tropical Zool., 14: 300.

Common Names

Scheffler's Puddle Frog (Channing and Howell, 2006, Amph. E. Afr.: 292).

Distribution

Semi-humid eastern Africa from central and southeastern Kenya and Tanzania (including Zanzibar) northwest up to central Uganda and eastern Rwanda, from sea level up to at least 1650 m

Geographic Occurrence

Natural Resident: Kenya, Rwanda, Tanzania, Uganda

Comment

Removed from the synonymy of Phrynobatrachus minutus by Largen, 2001, Tropical Zool., 14: 287–306, where it had been placed by Loveridge, 1929, Bull. U.S. Natl. Mus., 151: 107; Loveridge, 1936, Bull. Mus. Comp. Zool., 79: 421; Parker, Moreau, and Pakenham, 1940, Ann. Mag. Nat. Hist., Ser. 11, 5: 309–314. Channing and Howell, 2006, Amph. E. Afr.: 292–293, provided an account and noted that this species had previously been confused with Phrynobatrachus minutus. Pickersgill, 2007, Frog Search: 187–192, provided an account. Frétey, 2008, Alytes, 25: 99–172, summarized the literature. Schick, Zimkus, Channing, Köhler, and Lötters, 2010, Salamandra, 46: 24–36, discussed the species and provided an account. In species group B of Zimkus, Rödel, and Hillers, 2010, Mol. Phylogenet. Evol., 55: 883–900. Measey, Malonza, and Muchai, 2009, Amph. Taita Hills: 46–47, provided a brief account and photo for the Taita Hills, Kenya. Harper, Measey, Patrick, Menegon, and Vonesh, 2010, Field Guide Amph. E. Arc Mts. Tanzania and Kenya: 244–245, provided a brief account and photograph. Spawls, Wasonga, and Drewes, 2019, Amph. Kenya: 47, provided a photograph, range map, and brief characterization. In the Phrynobatrachus calcaratus group of Goutte, Reyes-Velasco, and Boissinot, 2019, ZooKeys, 824: 53–70. Channing and Rödel, 2019, Field Guide Frogs & Other Amph. Afr.: 310–311, provided a brief account, photograph, and range map. Dehling and Sinsch, 2023, Diversity, 15 (512): 1–81, discussed the range, identification, natural history, advertisement call, and conservation status in Rwanda.

External links:

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For access to general information see Wikipedia
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For access to relevant technical literature search Google Scholar
For images search CalPhoto Images and Google Images
To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist