Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Phrynobatrachus versicolor Ahl, 1924

Class: Amphibia > Order: Anura > Family: Phrynobatrachidae > Genus: Phrynobatrachus > Species: Phrynobatrachus versicolor

Phrynobatrachus versicolor Ahl, 1924, Zool. Anz., 61: 100. Syntypes: ZMB (215 specimens), unnumbered in the original publication; including ZMB 25370–71, 25423 (according to Deckert, 1938, Sitzungsber. Ges. Naturforsch. Freunde Berlin, 1938: 133), BMNH 1947.2.4.26 (formerly 1935.2.8.1, according to museum records), MCZ 17532–33 (on exchange from ZMB, according to Barbour and Loveridge, 1946, Bull. Mus. Comp. Zool., 96: 170), according to Grandison, 1985, in Frost (ed.), Amph. Species World: 451. Type localities: "Rugegewald", "Bugoiewald", and "Urwald nordwestlich des Tanganyika-Sees", Rwanda.

Phrynobatrachus (Natalobatrachus) versicolor — Laurent, 1941, Rev. Zool. Bot. Afr., 34: 196.

Phrynodon versicolor — Dubois, Ohler, and Pyron, 2021, Megataxa, 5: 419.

Common Names

Rwanda River Frog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 104).

Green Puddle Frog (Channing and Howell, 2006, Amph. E. Afr.: 297).

Distribution

Western Rwanda, northwestern Burundi, southwestern Uganda, and adjacent eastern Dem. Rep. Congo (North Kivu) in mountain swamps, 1500–2800 m elevation.

Geographic Occurrence

Natural Resident: Burundi, Congo, Democratic Republic of the, Rwanda, Uganda

Comment

See Laurent, 1972, Explor. Parc Natl. Virunga, Ser. 2, 22: 106, for discussion. Removed from the synonymy of Phrynobatrachus dendrobates by Laurent, 1941, Rev. Zool. Bot. Afr., 34: 196, where it had been placed by Loveridge, 1936, Field Mus. Nat. Hist. Publ., Zool. Ser., 22: 97. Channing and Howell, 2006, Amph. E. Afr.: 297–298, provided an account. Frétey, 2008, Alytes, 25: 99–172, summarized the literature. See photograph, map, description of geographic range and habitat, and conservation status in Stuart, Hoffmann, Chanson, Cox, Berridge, Ramani, and Young, 2008, Threatened Amph. World: 473. In species group A of Zimkus, Rödel, and Hillers, 2010, Mol. Phylogenet. Evol., 55: 883–900. Channing and Rödel, 2019, Field Guide Frogs & Other Amph. Afr.: 272–273, provided a brief account, photograph, and range map. In the Phrynobatrachus krefftii group of Goutte, Reyes-Velasco, and Boissinot, 2019, ZooKeys, 824: 53–70. Dehling and Sinsch, 2023, Diversity, 15 (512): 1–81, discussed the range, identification, natural history, advertisement call, and conservation status in Rwanda. Behangana, Byaruhanga, Magala, Katumba, Kagurusi, Dendi, and Luiselli, 2023, Wetlands, 43(88): 1–13, reported on presence in the wetlands of southwestern Uganda.

External links:

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For access to general information see Wikipedia
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To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.