Phrynobatrachus werneri (Nieden, 1910)

Class: Amphibia > Order: Anura > Family: Phrynobatrachidae > Genus: Phrynobatrachus > Species: Phrynobatrachus werneri

Arthroleptis werneri Nieden, 1910, Arch. Naturgesch., 76: 242. Syntypes: ZMB 20434, 20789 (total of 5 specimens), according to Grandison, 1985, in Frost (ed.), Amph. Species World: 451; ZMB 20434 designated lectotype by Blackburn and Rödel, 2011, Herpetologica, 67: 282. Type locality: "Banjobezirk" (= vicinity of Banyo) and "Bamenda", Cameroon; restricted to "Banjobezirk" (vicinity of Banyo) by lectotype designation.

Pseudarthroleptis werneriDeckert, 1938, Sitzungsber. Ges. Naturforsch. Freunde Berlin, 1938: 159.

Phrynobatrachus werneriLaurent, 1941, Rev. Zool. Bot. Afr., 34: 207.

Phrynobatrachus werneri werneriPerret, 1959, Bull. Soc. Neuchatel. Sci. Nat., 82: 251.

English Names

Werner's River Frog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 104).

Werner's Puddle Frog (Channing and Rödel, 2019, Field Guide Frogs & Other Amph. Afr.: 282).


Mount Kupe to Banyo, western Cameroon, mostly above 1100 m elevation. 


Schiøtz, 1963, Vidensk. Medd. Dansk Naturhist. Foren., 125: 36–38, provided records for Nigeria. Frétey, 2008, Alytes, 25: 99–172, summarized the literature. See comment under Phrynobatrachus manengoubensis, which may be a synonym, or Phrynobatrachus werneri may constitute more than one species according to Zimkus, 2009, Zool. J. Linn. Soc., 157: 796–813 (referring to what was subsequently named as Phrynobatrachus schioetzi). In species group C of Zimkus, Rödel, and Hillers, 2010, Mol. Phylogenet. Evol., 55: 883–900, whose analysis suggested that nominal Phrynobatrachus werneri is a species complex forming a paraphyletic series with respect to Phrynobatrachus manengoubensis. Blackburn and Rödel, 2011, Herpetologica, 67: 282, discussed the realignment of range with the description of Phrynobatrachus schioetziChanning and Rödel, 2019, Field Guide Frogs & Other Amph. Afr.: 282–283, provided a brief account, photograph, and range map. In the Phrynobatrachus natalensis group of Goutte, Reyes-Velasco, and Boissinot, 2019, ZooKeys, 824: 53–70. In the Phrynobatracus werneri group according to Gvoždík, Nečas, Dolinay, Zimkus, Schmitz, and Fokam, 2020, PeerJ, 8 (e8393): 1–52.  

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