Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Ptychadena grandisonae Laurent, 1954

Class: Amphibia > Order: Anura > Family: Ptychadenidae > Genus: Ptychadena > Species: Ptychadena grandisonae

Ptychadena grandisonae Laurent, 1954, Ann. Mus. R. Congo Belge, Tervuren, Zool., 34: 11. Holotype: MD 506, by original designation; now MRAC B-60530, according to Ceríaco, Marques, André, Afonso, Blackburn, and Bauer, 2020, Bull. Mus. Comp. Zool., 162: 410. Type locality: "Muita [River], Luembe E., [Lunda Sul Province,] Angola"; given as "'Canzar, Muita' [Lunda Sul Province]", Angola, Lat. -7.80 Long. 21.45 by Ceríaco, Marques, André, Afonso, Blackburn, and Bauer, 2020, Bull. Mus. Comp. Zool., 162: 409.

Rana (Ptychadena) grandisonae — Guibé and Lamotte, 1957, Bull. Inst. Franç. Afr. Noire, Ser. A, 19: 942, by implication.

Rana grandisonae — Schmidt and Inger, 1959, Explor. Parc Natl. Upemba, Miss. G.F. de Witte, 56: 70–75.

Ptychadena grandisonae — Poynton, 1970, Ann. Natal Mus., 20: 370; Dubois, 1987 "1986", Alytes, 5: 56; by implication.

Rana (Ptychadena) grandisonae — Dubois, 1981, Monit. Zool. Ital., N.S., Suppl., 15: 233, by implication.

Ptychadena (Ptychadena) grandisonae — Dubois, 1992, Bull. Mens. Soc. Linn. Lyon, 61: 316.

Common Names

Muita Grassland Frog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 105).

Grandison's Ridged Frog (Channing, 2001, Amph. Cent. S. Afr.: 325).

Grandison's Grass Frog (Channing and Rödel, 2019, Field Guide Frogs & Other Amph. Afr.: 338).

Many-ridged Grass Frog (Conradie, Keates, Verburgt, Baptista, and Harvey, 2023, Amph. Rept. Conserv., 17: 43).

Distribution

Open savannas in southwestern to northeastern Angola through southern Dem. Rep. Congo to Burundi to western Tanzania, and northern and western Zambia.

Geographic Occurrence

Natural Resident: Angola, Burundi, Congo, Democratic Republic of the, Tanzania, Zambia

Comment

Reviewed by Schmidt and Inger, 1959, Explor. Parc Natl. Upemba, Miss. G.F. de Witte, 56: 70-75. Specimens cited by Amiet, 1973, Ann. Fac. Sci. Cameroun, 13: 137, and Amiet, 1974, Ann. Fac. Sci. Cameroun, 18: 112, were confused with Ptychadena straeleni, according to Perret, 1979, Bull. Soc. Neuchatel. Sci. Nat., 102: 18. See also accounts by Poynton and Broadley, 1985, Ann. Natal Mus., 27: 150, and Channing, 2001, Amph. Cent. S. Afr.: 325. Harper, Measey, Patrick, Menegon, and Vonesh, 2010, Field Guide Amph. E. Arc Mts. Tanzania and Kenya: 262–263, provided a brief account and photograph. Marques, Ceríaco, Blackburn, and Bauer, 2018, Proc. California Acad. Sci., Ser. 4, 65 (Suppl. II): 132–133, provided a map for Angola and brief discussion of literature.See Bittencourt-Silva, 2019, Amph. Rept. Conserv., 13 (2: e181): 11, for comment on population in western Zambia. Channing and Rödel, 2019, Field Guide Frogs & Other Amph. Afr.: 338–339, provided a brief account, photograph, and range map. Dehling and Sinsch, 2023, Diversity, 15 (512): 67, discussed erroneous records for Rwanda. Conradie, Keates, Verburgt, Baptista, and Harvey, 2023, Amph. Rept. Conserv., 17: 43, provided an account and dot map of the species in Angola.

External links:

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For access to general information see Wikipedia
For additional sources of general information from other websites search Google
For access to relevant technical literature search Google Scholar
For images search CalPhoto Images and Google Images
To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.