Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Ptychadena neumanni (Ahl, 1924)

Class: Amphibia > Order: Anura > Family: Ptychadenidae > Genus: Ptychadena > Species: Ptychadena neumanni

Rana neumanni Ahl, 1924, Mitt. Zool. Mus. Berlin, 11: 4. Syntypes: 35 syntypes noted in original publication; apparently including ZMB 26879 (3 specimens), and MCZ 22331-32 (according to Barbour and Loveridge, 1946, Bull. Mus. Comp. Zool., 96: 184). Restricted to ZMB 26879 (3 specimens) by inference of restricted type locality and action of Largen, 2001, Tropical Zool., 14: 340. Type locality: "Didda", "Somaliland", "Gadat (Gofa)", and "Ufa"; restricted to "Gadat (Gofa), Sud Ethiopia" by Perret, 1980, Monit. Zool. Ital., N.S., Suppl., 13: 157. Largen, 2001, Tropical Zool., 14: 340 noted that ZMB 26877 and 26878A-T are referrable to Ptychadena nana.

Ptychadena neumanni — Perret, 1980, Monit. Zool. Ital., N.S., Suppl., 13: 157; Dubois, 1987 "1986", Alytes, 5: 56; by implication

Rana (Ptychadena) neumanni — Dubois, 1981, Monit. Zool. Ital., N.S., Suppl., 15: 233, by implication.

Ptychadena (Ptychadena) neumanni — Dubois, 1992, Bull. Mens. Soc. Linn. Lyon, 61: 316.

Ptychadena largeni Perret, 1994, Bull. Soc. Neuchatel. Sci. Nat., 117: 67. Holotype: MHNG 2513.31, by original designation. Type locality: "Addis Ababa, Shewa (Shoa) 09° 02 N 38° 45 E, alt. 2500 m", Ethiopia. Synonymy by Largen, 2001, Tropical Zool., 14: 340.

Ptychadena nuerensis Smith, Noonan, and Colston, 2017, R. Soc. Open Sci., 4(170021): 11. Holotype: Timothy J. Colston 455, by original designation. Type locality: "Telouse, Lare, Gambela Region, Ethiopia, 8.26875° N, 33.94616° E, 415 m". Invalid nomenclatural act due to publishing a new name in an electronic journal without zoobank number.

English Names

Neumann's Grassland Frog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 105).

Neumann's Grass Frog (Largen and Spawls, 2010, Amph. Rept. Ethiopia Eritrea: 189).

Distribution

Ethiopian Plateau, 820-3800 m elevation. See comment below regarding the ranges of the various unnamed lineages within this complex.

Comment

See comment under Ptychadena nana. Largen, 2001, Tropical Zool., 14: 338–339, discussed distribution and the difficulty in separating Ptychadena erlangeri and Ptychadena neumanni by external morphology alone. See discussion by Largen, 2001, Tropical Zool., 14: 340–342. See account, photograph, and map for Ethiopia by Largen and Spawls, 2010, Amph. Rept. Ethiopia Eritrea: 188–190, who suggested that this name represents a complex of species of which Ptychadena largeni is only questionably separated. Freilich, Tollis, and Boissinot, 2014, Mol. Phylogenet. Evol., 71: 157–169, presented molecular evidence that nominal Ptychadena neumanni is composed of at least five species, two of which (sp. 2 and 3) form the sister taxon of Ptychadena erlangeri, one of which is the sister taxon of that group (sp. 4), another that is the sister taxon of that inclusive group (sp. 1), and a fifth (sp. 5) that is the sister taxon of Ptychadena cooperi. This overall complex surrounds Ptychadena nana phylogenetically as well. Geographically they are known definitively from: 1: moist evergreen forests of the southwest to the grasslands in the vicinity of Addis Ababa; 2: widely distributed on both side of the Great Rift and limited to grasslands (Ptychadena largeni); 3: Bale Mountains (now Ptychadena levenorum); 4: southern part of the Eastern Highlands (now Ptychadena goweri); 5: western highlands, north of the Blue Nile Valley (now Ptychadena amharensis). Channing and Rödel, 2019, Field Guide Frogs & Other Amph. Afr.: 340–341, provided a brief account, photograph, and range map.

External links:

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For additional sources of information from other sites search Google
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To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For related information on conservation and images as well as observation see iNaturalist; for a quick link to their maps see iNaturalist KML
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.