Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Ptychadena upembae (Schmidt and Inger, 1959)

Class: Amphibia > Order: Anura > Family: Ptychadenidae > Genus: Ptychadena > Species: Ptychadena upembae

Rana (Ptychadena) upembae Schmidt and Inger, 1959, Explor. Parc Natl. Upemba, Miss. G.F. de Witte, 56: 111. Holotype: MRAC (formerly Inst. Parcs Natl. Congo Belge) 1228 according to Lang, 1990, Doc. Trav., Inst. R. Sci. Nat. Belg., 59: 13. Type locality: "Kaswabilenga, Parc National de l'Upemba, Province Katanga, Belgian Congo [=Dem. Rep. Congo]".

Ptychadena upembae — Poynton, 1964, Senckenb. Biol., 45: 207; Dubois, 1987 "1986", Alytes, 5: 56; by implication.

Ptychadena upembae machadoi Laurent, 1964, Publ. Cult. Companhia Diamantes Angola, 67: 134. Holotype: MD 5364, by original designation. Type locality: "Alto Chicapa, Lunda [Sur Province]", Angola.

Ptychadena upembae upembae — Laurent, 1964, Publ. Cult. Companhia Diamantes Angola, 67: 134, by implication.

Rana (Ptychadena) upembae upembae — Dubois, 1981, Monit. Zool. Ital., N.S., Suppl., 15: 233, by implication.

Rana (Ptychadena) upembae machadoi — Dubois, 1981, Monit. Zool. Ital., N.S., Suppl., 15: 233, by implication.

Ptychadena (Ptychadena) upembae — Dubois, 1992, Bull. Mens. Soc. Linn. Lyon, 61: 316.

Common Names

Upemba Grassland Frog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 106).

Upemba Ridged Frog (Ananjeva, Borkin, Darevsky, and Orlov, 1988, Dict. Amph. Rept. Five Languages: 129; Channing, 2001, Amph. Cent. S. Afr.: 342–344; Channing and Howell, 2006, Amph. E. Afr.: 316).

Upemba Grass Frog (Channing and Rödel, 2019, Field Guide Frogs & Other Amph. Afr.: 338).

Distribution

Northwestern to southeastern and northeastern Angola, Katanga Province (Dem. Rep. Congo), Zambia, Malawi, and northern Mozambique; possibly into southwestern Tanzania.

Geographic Occurrence

Natural Resident: Angola, Congo, Democratic Republic of the, Malawi, Mozambique, Zambia

Likely/Controversially Present: Tanzania

Comment

Discussed by Stewart, 1967, Amph. Malawi: 76–77. See Laurent, 1964, Publ. Cult. Companhia Diamantes Angola, 67: 134, for a subspecies (Ptychadena upembae machadoi). See also Poynton and Broadley, 1985, Ann. Natal Mus., 27: 150–151, fand Channing, 2001, Amph. Cent. S. Afr.: 342–344, for accounts of southern population. Channing and Howell, 2006, Amph. E. Afr.: 316–317, provided an account for East Africa. Mercurio, 2011, Amph. Malawi: 271–273, provided an account for Malawi. Marques, Ceríaco, Blackburn, and Bauer, 2018, Proc. California Acad. Sci., Ser. 4, 65 (Suppl. II): 142, provided a map for Angola and brief discussion of the literature. See Bittencourt-Silva, 2019, Amph. Rept. Conserv., 13 (2: e181): 12, for comment on population in western Zambia. Channing and Rödel, 2019, Field Guide Frogs & Other Amph. Afr.: 338–339, provided a brief account, photograph, and range map. Ceríaco, Marques, André, Afonso, Blackburn, and Bauer, 2020, Bull. Mus. Comp. Zool., 162: 412–413, discussed and pictured the holotype and the dispersal of paratypes of Ptychadena upembae machadoi. Conradie, Keates, Verburgt, Baptista, and Harvey, 2023, Amph. Rept. Conserv., 17: 43–44, provided an account and dot map of the species in Angola.

External links:

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For access to general information see Wikipedia
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For images search CalPhoto Images and Google Images
To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.