Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Aquarana heckscheri (Wright, 1924)

Class: Amphibia > Order: Anura > Family: Ranidae > Genus: Aquarana > Species: Aquarana heckscheri

Rana heckscheri Wright, 1924, Proc. Biol. Soc. Washington, 37: 143. Holotype: CUMV 1025; apparently lost, although Figs. 5 and 6 in Plate 38 of Wright, 1932, Life Hist. Frogs Okefinokee Swamp, 2, are of the holotype, according to Sanders, 1984, Cat. Am. Amph. Rept., 348: 2. Type locality: "Alligator Swamp, Callahan, [Nassau County,] Florida", USA.

Rana (Rana) heckscheri — Dubois, 1987 "1986", Alytes, 5: 41–42, by implication.

Rana (Aquarana) heckscheri — Dubois, 1992, Bull. Mens. Soc. Linn. Lyon, 61: 331.

Rana (Novirana, Aquarana) heckscheri — Hillis and Wilcox, 2005, Mol. Phylogenet. Evol., 34: 305. See Dubois, 2006, Mol. Phylogenet. Evol., 42: 317–330, Hillis, 2007, Mol. Phylogenet. Evol., 42: 331–338, and Dubois, 2007, Cladistics, 23: 390–402, for relevant discussion of nomenclature. Invalid name formulation under the International Code of Zoological Nomenclature (1999) as discussed by Dubois, 2007, Cladistics, 23: 395.

Rana (Novirana) heckscheri — Hillis and Wilcox, 2005, Mol. Phylogenet. Evol., 34: 305. Nomenclatural act of Hillis and Wilcox, 2005, as interpreted by Fouquette and Dubois, 2014, Checklist N.A. Amph. Rept.: 417.

Lithobates heckscheri — Frost, Grant, Faivovich, Bain, Haas, Haddad, de Sá, Channing, Wilkinson, Donnellan, Raxworthy, Campbell, Blotto, Moler, Drewes, Nussbaum, Lynch, Green, and Wheeler, 2006, Bull. Am. Mus. Nat. Hist., 297: 369; Che, Pang, Zhao, Wu, Zhao, and Zhang, 2007, Mol. Phylogenet. Evol., 43: 1–13; by implication.

Lithobates (Aquarana) heckscheri — Dubois, 2006, C. R. Biol., Paris, 329: 829; Dubois, 2006, Mol. Phylogenet. Evol., 42: 325.

Rana (Aquarana) heckscheri — Hillis, 2007, Mol. Phylogenet. Evol., 42: 335–336, by implication and combination made only for discussion; Yuan, Zhou, Chen, Poyarkov, Chen, Jang-Liaw, Chou, Matzke, Iizuka, Min, Kuzmin, Zhang, Cannatella, Hillis, and Che, 2016, Syst. Biol., 65: 835.

Rana (Lithobates) heckscheri — Fouquette and Dubois, 2014, Checklist N.A. Amph. Rept.: 417.

Aquarana heckscheri — Dubois, Ohler, and Pyron, 2021, Megataxa, 5: 424.

Common Names

Heckscher's Frog (Wright, 1932, Life Hist. Frogs Okefinokee Swamp, 2: 14).

River-Swamp Frog (Wright and Wright, 1933, Handb. Frogs Toads U.S. Canada: xi; Carr, 1940, Univ. Florida Biol. Sci. Ser., 3: 66).

River Swamp Bullfrog (Schmidt, 1953, Check List N. Am. Amph. Rept., Ed. 6: 80).

River Frog (Conant, Cagle, Goin, Lowe, Neill, Netting, Schmidt, Shaw, Stebbins, and Bogert, 1956, Copeia, 1956: 177; Conant, 1975, Field Guide Rept. Amph. E. Cent. N. Am., Ed. 2: 339; Collins, Huheey, Knight, and Smith, 1978, Herpetol. Circ., 7: 13; Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 107; Collins, 1997, Herpetol. Circ., 25: 13; Crother, Boundy, Campbell, de Queiroz, Frost, Highton, Iverson, Meylan, Reeder, Seidel, Sites, Taggart, Tilley, and Wake, 2001 "2000", Herpetol. Circ., 29: 14; Frost, McDiarmid, and Mendelson, 2008, in Crother (ed.), Herpetol. Circ., 37: 8; Collins and Taggart, 2009, Standard Common Curr. Sci. Names N. Am. Amph. Turtles Rept. Crocodil., ed. 6: 8; Frost, McDiarmid, Mendelson, and Green, 2012, in Crother (ed.), Herpetol. Circ., 39: 18; Frost, Lemmon, McDiarmid, and Mendelson, 2017, in Crother (ed.), Herpetol. Circ., 43: 15).

Distribution

Southern North Carolina and southern Mississippi to northern Florida on the coastal plain, USA; introduced into China.

Geographic Occurrence

Natural Resident: United States of America, United States of America - Alabama, United States of America - Florida, United States of America - Georgia, United States of America - Mississippi, United States of America - North Carolina, United States of America - South Carolina

Endemic: United States of America

Introduced: China, People's Republic of

Comment

In the Rana catesbeiana group of XXX. In the equivalent Rana clamitans group of Dubois, 1987 "1986", Alytes, 5: 41, and in the equivalent subgenus Aquarana of Dubois, 1992, Bull. Mens. Soc. Linn. Lyon, 61: 331. Reviewed by Sanders, 1984, Cat. Am. Amph. Rept., 348: 1–2. Fei, 1999, Atlas Amph. China: 308–309, provided a brief account and figure for the species in China. Fei and Ye, 2001, Color Handbook Amph. Sichuan: 230, provided a brief account and illustration for the introduced Chinese population. Butterfield and Lannoo, 2005, in Lannoo (ed.), Amph. Declines: 557–558, and Dodd, 2013, Frogs U.S. and Canada, 2 : 556–560, provided accounts that summarized relevant literature of biology, range, and conservation. Elliot, Gerhardt, and Davidson, 2009, Frogs and Toads of N. Am.: 206–207, provided an account, photos, and advertisement call. Guyer and Bailey, 2023, Frogs and Toads of Alabama: 125–127, provided a detailed account for Alabama, USA.

External links:

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For access to general information see Wikipedia
For additional sources of general information from other websites search Google
For access to relevant technical literature search Google Scholar
For images search CalPhoto Images and Google Images
To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist
For additional information specific to China see Amphibia China
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.