Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Chalcorana labialis (Boulenger, 1887)

Class: Amphibia > Order: Anura > Family: Ranidae > Genus: Chalcorana > Species: Chalcorana labialis

Rana labialis Boulenger, 1887, Ann. Mag. Nat. Hist., Ser. 5, 19: 345. Syntypes: BMNH ("several" specimens), by original designation. BMNH 1947.2.3.40 designated lectotype by Inger, Stuart, and Iskandar, 2009, Zool. J. Linn. Soc., 155: 132. Type locality: "within a radius of fifty miles from the town of Malacca", West Malaysia.

Rana (Hylorana) chalconota var. labialis — Boulenger, 1920, Rec. Indian Mus., 20: 203.

Rana (Hylarana) labialis — Van Kampen, 1923, Amph. Indo-Austral. Arch.: 220.

Hydrophylax labialis — Frost, 2007, Amph. Spec. World, vers. 5.0, new combination; by implication of results of Frost, Grant, Faivovich, Bain, Haas, Haddad, de Sá, Channing, Wilkinson, Donnellan, Raxworthy, Campbell, Blotto, Moler, Drewes, Nussbaum, Lynch, Green, and Wheeler, 2006, Bull. Am. Mus. Nat. Hist., 297: 368.

Hylarana labialis — Che, Pang, Zhao, Wu, Zhao, and Zhang, 2007, Mol. Phylogenet. Evol., 43: 1-13, by implication; Dubois, Ohler, and Pyron, 2021, Megataxa, 5: 423.

Chalcorana labialis — Fei, Ye, and Jiang, 2010, Herpetol. Sinica, 12: 33; Oliver, Prendini, Kraus, and Raxworthy, 2015, Mol. Phylogenet. Evol., 90: 187.

English Names

White-lipped Frog (Chan-ard, 2003, Photograph. Guide Amph. Thailand: 134 [as Rana raniceps; Dinesh, Radhakrishnan, Gururaja, and Bhatta, 2009, Rec. Zool. Surv. India, Occas. Pap., 302: 75; Grismer, 2012, Field Guide Amph. Rept. Seribuat Arch.: 74).

Distribution

West Malaysia; possibly Singapore.

Comment

Removed from the synonymy of Hylarana raniceps by Stuart, Inger, and Voris, 2006, Biol. Lett., 2: 470-474, where it had been placed by Inger, 1966, Fieldiana, Zool., 52: 177. Stuart, Inger, and Voris, 2006, Biol. Lett., 2: 470-474, also noted that this species is morphologically indistinguishable from Hylarana chalconota, and did not detail the range, however, which is highly provisional. Inger, Stuart, and Iskandar, 2009, Zool. J. Linn. Soc., 155: 123-147, redelimited the species and discussed its phylogenetic placement with respect to other members of the Hylarana chalconota group, and noted an unnamed species that co-occurs in West Malaysia. See brief account by Grismer, 2012, Field Guide Amph. Rept. Seribuat Arch.: 74-75, for the Seribuat Archipelago, West Malaysia. Shahriza and Ibrahim, 2014, Check List, 10: 253–259, provided a photograph and brief natural history observations for a population in Kedah, West Malaysia. See comments on morphology and natural history in West Malaysia by Chan, Muin, Anuar, Andam, Razak, and Aziz, 2019, Check List, 15: 1055–1069. Hong, Anuar, Grismer, and Quah, 2021, Check List, 17: 797, reported the species from Batu Hampar Recreational Forest, Kedah, West Malaysia.

External links:

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