Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Hylarana darlingi (Boulenger, 1902)

Class: Amphibia > Order: Anura > Family: Ranidae > Genus: Hylarana > Species: Hylarana darlingi

Rana darlingi Boulenger, 1902, Proc. Zool. Soc. London, 1902: 15. Syntypes: BMNH (2 specimens), by original designation; these being BMNH 1947.2.2.67–68 (originally numbered 1902.2.12.114-115), by museum records. Type locality: "Mazöe and between Umtali [Mutare] and Marandellas [= Marondera]", Mashonaland, Zimbabwe.

Rana (Rana) darlingi — Monard, 1937 "1936", Bull. Soc. Neuchatel. Sci. Nat., 62: 44.

Hylarana darlingi — Poynton, 1964, Ann. Natal Mus., 17: 119.

Rana (Hylarana) darlingi — Dubois, 1987 "1986", Alytes, 5: 42, by implication.

Rana (Amnirana) darlingi — Dubois, 1992, Bull. Mens. Soc. Linn. Lyon, 61: 324.

Amnirana darlingi — Channing, 2001, Amph. Cent. S. Afr.: 266; Fei, Ye, and Jiang, 2010, Herpetol. Sinica, 12: 33. See comment under Ranidae record.

Hydrophylax darlingi — Frost, Grant, Faivovich, Bain, Haas, Haddad, de Sá, Channing, Wilkinson, Donnellan, Raxworthy, Campbell, Blotto, Moler, Drewes, Nussbaum, Lynch, Green, and Wheeler, 2006, Bull. Am. Mus. Nat. Hist., 297: 368.

Hylarana darlingi — Che, Pang, Zhao, Wu, Zhao, and Zhang, 2007, Mol. Phylogenet. Evol., 43: 1-13, by implication; Dubois, Ohler, and Pyron, 2021, Megataxa, 5: 461, by implication.

Amnirana darlingi — Oliver, Prendini, Kraus, and Raxworthy, 2015, Mol. Phylogenet. Evol., 90: 187.

English Names

Golden-backed Frog (Broadley, 1971, Puku, 6: 113; Broadley, 1973, J. Herpetol. Assoc. Afr., 10: 23; Lambiris, 1990 "1989", Monogr. Mus. Reg. Sci. Nat. Torino, 10: 100).

Darling's Frog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 107).

Darling's Golden-backed Frog (Van Dijk, 1978, J. Herpetol. Assoc. Afr., 17: 15; Du Preez and Carruthers, 2009, Compl. Guide Frogs S. Afr.: 452).

Darling's White-lipped Frog (Channing, 2001, Amph. Cent. S. Afr.: 266).

Distribution

Savannas of Katanga Province (Dem. Rep. Congo), extreme northern Botswana and adjacent Namibia, Zimbabwe, southern Malawi, and western Mozambique; reported from northeastern Dem. Rep. Congo.

Geographic Occurrence

Natural Resident: Botswana, Congo, Democratic Republic of the, Malawi, Mozambique, Namibia, Zimbabwe

Comment

Records from Angola and western Zambia presumably apply to Hylarana adiscifera, recognized as distinct in 2023. Sole member of the Rana (Amnirana) darlingi group of Dubois, 1992, Bull. Mens. Soc. Linn. Lyon, 61: 324. See Poynton, 1964, Ann. Natal Mus., 17: 119, and Perret, 1977, Rev. Suisse Zool., 84: 843. See also Laurent, 1964, Publ. Cult. Companhia Diamantes Angola, 67: 132. See accounts by Poynton and Broadley, 1985, Ann. Natal Mus., 27: 139-140 (as Hylarana darlingi). Channing, 2001, Amph. Cent. S. Afr.: 266-268, and Pickersgill, 2007, Frog Search: 87-88 (as Amnirana darlingi), and Du Preez and Carruthers, 2009, Compl. Guide Frogs S. Afr.: 452-453. Mercurio, 2011, Amph. Malawi: 298-299, provided an account for Malawi. Conradie, Tweddle, Mäkinen, and Bills, 2012, Zootaxa, 3441: 56-58. reported on larval morphology. Channing, Rödel, and Channing, 2012, Tadpoles of Africa: 364, provided information on comparative larval morphology. See range map provided by Jongsma, Barej, Barratt, Burger, Conradie, Ernst, Greenbaum, Hirschfeld, Leaché, Penner, Portik, Zassi-Boulou, Rödel, and Blackburn, 2018, Mol. Phylogenet. Evol., 120: 275. Marques, Ceríaco, Blackburn, and Bauer, 2018, Proc. California Acad. Sci., Ser. 4, 65 (Suppl. II): 158–160, provided a map for Angola and brief discussion of the literature. Mali, Banda, Chifundera, Badjedjea, Sebe, Lokasola, Ewango, Tungaluna, and Akaibe, 2019, Am. J. Zool., 2: 38–43, reported the species from the Okapi Wildlife Reserve, northeastern Dem. Rep. Congo, but this identification requires confirmation. Channing and Rödel, 2019, Field Guide Frogs & Other Amph. Afr.: 388–389, provided a brief account, photograph, and range map.

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