Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Hylarana intermedia (Rao, 1937)

Class: Amphibia > Order: Anura > Family: Ranidae > Genus: Hylarana > Species: Hylarana intermedia

Rana (Hylorana) intermedius Rao, 1937, Proc. Indian Acad. Sci., Ser. B, 6: 394. Holotype: CCB; now lost according to Dubois, 1984, Alytes, 3: 157. BNHS 5823, from type locality, designated neotype by Biju, Garg, Mahony, Wijayathilaka, Senevirathne, and Meegaskumbura, 2014, Contr. Zool., Amsterdam, 83: 295. Type locality: "Saklespur, Hassan District, Mysore [now Karnataka] State", India. Incorrect gender of species name.

Sylvirana intermedia — Das and Dutta, 2007, Hamadryad, 31: 164.

Hylarana intermedius — Biju, Garg, Mahony, Wijayathilaka, Senevirathne, and Meegaskumbura, 2014, Contr. Zool., Amsterdam, 83: 295. Incorrect gender of species name.

Indosylvirana intermedia — Oliver, Prendini, Kraus, and Raxworthy, 2015, Mol. Phylogenet. Evol., 90: 190.

Hylarana intermedia — Dubois, Ohler, and Pyron, 2021, Megataxa, 5: 501, by implication.

English Names

Rao's Intermediate Golden-backed Frog (Biju, Garg, Mahony, Wijayathilaka, Senevirathne, and Meegaskumbura, 2014, Contr. Zool., Amsterdam, 83: 295).

Rao's Brown-coloured Frog (Chanda, 2002, Handb. Indian Amph.: 115).

Distribution

Western Ghats north of the Palghat Gap of the states of Karnataka and adjoining regions of Kerala, predominently at elevations between 600 and 1183 m, but also as low as 8 m elevation at Adyar (Mangalore).

Geographic Occurrence

Natural Resident: India

Endemic: India

Comment

Removed from the synonymy of Hylarana flavescens by Biju, Garg, Mahony, Wijayathilaka, Senevirathne, and Meegaskumbura, 2014, Contr. Zool., Amsterdam, 83: 295 (who provided an account and placed it in their Hylarana aurantiaca group), where it had been placed by Dubois, 1992, Bull. Mens. Soc. Linn. Lyon, 61: 341. Chanda, 2002, Handb. Indian Amph.: 115, provided a brief account (as Rana intermedius). Das and Dutta, 2007, Hamadryad, 31: 154–181, noted no larval descriptions in the literature. Kamath and Sreekar, 2016, Acta Herpetol., Firenze, 11: 15–20, reported on morphology, ecology, and behavior.

External links:

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To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observation see iNaturalist
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.