Hylarana milleti (Smith, 1921)

Class: Amphibia > Order: Anura > Family: Ranidae > Genus: Hylarana > Species: Hylarana milleti

Rana milleti Smith, 1921, Proc. Zool. Soc. London, 1921: 431. Syntypes: BMNH1947.2.1.37–46 (formerly numbered 1921.4.1.254–263), by museum records (formerly M. Smith 5119, 5128–29, 2571, 2600, 2602, 4818, 5103, 5107, 5176); including a specimen in the FMNH according to Marx, 1976, Fieldiana, Zool., 69: 63. Type locality: "Dalat" (9 specimens) and "Lang Bian" (M. Smith 5176), Langbian Plateau, southern Annam, Vietnam.

Hylarana milleti — Bourret, 1939, Annexe Bull. Gen. Instr. Publique, Hanoi, 1939: 59.

Rana (Hylarana) milleti — Bourret, 1942, Batr. Indochine: 312; Dubois, 1987 "1986", Alytes, 5: 42, by implication.

Rana (Sylvirana) milleti — Dubois, 1992, Bull. Mens. Soc. Linn. Lyon, 61: 326.

Hylarana milleti — Chen, Murphy, Lathrop, Ngo, Orlov, Ho, and Somorjai, 2005, Herpetol. J., 15: 237, by implication; Che, Pang, Zhao, Wu, Zhao, and Zhang, 2007, Mol. Phylogenet. Evol., 43: 3, by implication; Dubois, Ohler, and Pyron, 2021, Megataxa, 5: 423.

Sylvirana milleti — Frost, Grant, Faivovich, Bain, Haas, Haddad, de Sá, Channing, Wilkinson, Donnellan, Raxworthy, Campbell, Blotto, Moler, Drewes, Nussbaum, Lynch, Green, and Wheeler, 2006, Bull. Am. Mus. Nat. Hist., 297: 370.

Hylarana (Sylvirana) milleti — Fei, Ye, Jiang, and Xie, 2008, Acta Zootaxon. Sinica, 33: 206.

Sylvirana milleti — Fei, Ye, and Jiang, 2010, Herpetol. Sinica, 12: 34. See comment under Ranidae.

Indosylvirana milleti — Oliver, Prendini, Kraus, and Raxworthy, 2015, Mol. Phylogenet. Evol., 90: 190. 

Papurana milleti — Chan, Hutter, Wood, Grismer, and Brown, 2020, Mol. Phylogenet. Evol., 151 (106899): 1. 

Hylarana (Pterorana) milleti — Wu, Xu, Suwannapoom, Nguyen, Murphy, Papenfuss, Lathrop, Kilunda, Gao, Yuan, Chen, Zhang, Zhao, Wang, Rahman, Nneji, Zhao, Wang, Jin, Zhang, and Che, 2024 "2025", Mol. Phylogenet. Evol., 108223: 1–11, by implication.

Common Names

Millet's Frog (Chan-ard, 2003, Photograph. Guide Amph. Thailand: 130).

Millet's Forestfrog (Zug and Mulcahy, 2020 "2019", Amph. Rept. S. Tanintharyi: 55; Zug, 2022, Smithson. Contrib. Zool., 653: 39). 

Dalat Frog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 108).

Dalat Forest Frog (Poyarkov, Nguyen, Popov, Geissler, Pawangkhanant, Neang, Suwannapoom, and Orlov, 2021, Russ. J. Herpetol., 28 (3A): 51). 

Distribution

Isolated localities in southern Vietnam and southern and eastern Thailand (Amnat Charoen, Nakhon Ratchasima, Chachoengsao, and Chantaburi provinces); southwestern Cambodia; Mo-han, Yunnan, on the China-Laos border; reported from South Tanintharyi, peninsular Myanmar; expected in northern and southern Laos.

Geographic Occurrence

Natural Resident: Cambodia, China, People's Republic of, Myanmar, Thailand, Vietnam

Likely/Controversially Present: Laos

Comment

Prior to the revision of Frost, Grant, Faivovich, Bain, Haas, Haddad, de Sá, Channing, Wilkinson, Donnellan, Raxworthy, Campbell, Blotto, Moler, Drewes, Nussbaum, Lynch, Green, and Wheeler, 2006, Bull. Am. Mus. Nat. Hist., 297, placed in the Section Hylarana, subsection Hydrophylax, subgenus Sylvirana of Dubois, 1992, Bull. Mens. Soc. Linn. Lyon, 61: 326. See account by Bourret, 1942, Batr. Indochine: 312–314. See comments by Inger, Orlov, and Darevsky, 1999, Fieldiana, Zool., N.S., 92: 20–21. Chan-ard, 2003, Photograph. Guide Amph. Thailand: 130–131, provided a very brief account, map for Thailand, and photograph. Nguyen, Ho, and Nguyen, 2005, Checklist Amph. Rept. Vietnam: 30, provided specific localities for Vietnam. Chuaynkern, Ohler, Inthara, Kumtong, and Dubois, 2004, Nat. Hist. J. Chulalongkorn Univ., 4: 1–13, discussed the range. Stuart and Emmett, 2006, Fieldiana, Zool., N.S., 109: 9, provided a record for the Cardamom Mountains, southwestern Cambodia. See discussion by Ohler, 2007, Alytes, 25: 55–74. Fei, Ye, Jiang, and Xie, 2008, Acta Zootaxon. Sinica, 33: 199–206, provided a key to the close relatives of this species. They distinguished Hylarana bannanica from Hylarana milleti, unlike Ohler, 2007, Alytes, 25: 70, who placed them in synonymy on the basis of morphological characteristics, which appeared after the paper by Fei et al. had gone to press. Neang and Holden, 2008, Field Guide Amph. Cambodia: 81, provided a photograph, brief account of identification, ecology, and range in Cambodia. Fei, Ye, and Jiang, 2010, Colored Atlas of Chinese Amph.: 302–303, provided a brief account (as Hylarana bannanica). Chan-ard, Cota, and Makchai, 2011, Amph. E. Region Thailand: 104–105, provided a photograph and brief account for the Eastern Region of Thailand, and (p. 145) detailed the range in Thailand. See account, photograph, and map for Vietnam in Vassilieva, Galoyan, Poyarkov, and Geissler, 2016, Photograph. Field Guide Amph. Rept. Lowland S. Vietnam: 104–105. Niyomwan, Srisom, and Pawangkhanant, 2019, Field Guide Amph. Thailand: 346–347, provided a brief account (description, photographs, habitat, and range) for Thailand (in Thai). Zug and Mulcahy, 2020 "2019", Amph. Rept. S. Tanintharyi: 55–56, provided a brief account for South Tanintharyi, peninsular Myanmar. Zug, 2022, Smithson. Contrib. Zool., 653: 39, briefly discussed identification, habitat, and range in Myanmar. 

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