Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Lithobates kauffeldi (Feinberg, Newman, Watkins-Colwell, Schlesinger, Zarate, Curry, Shaffer, and Burger, 2014)

Class: Amphibia > Order: Anura > Family: Ranidae > Genus: Lithobates > Species: Lithobates kauffeldi

Rana kauffeldi Feinberg, Newman, Watkins-Colwell, Schlesinger, Zarate, Curry, Shaffer, and Burger, 2014, PLoS One, 9 (10: e108213): 4. Holotype: YPM 13217, by original designation. Type locality: "Bloomfield region, Richmond County (Staten Island), N[ew] Y[ork], United States". urn:lsid:zoobank.org:act:149ED690-FA7D-4216-A6A1-AA48CC39B292.

Lithobates kauffeldi — Frost, 2014, Amph. Spec. World, vers. 6.0 (of 31 Oct.). To bring the taxonomy into alignment with that employed by this online catalogue.

Rana (Pantherana) kauffeldi — Yuan, Zhou, Chen, Poyarkov, Chen, Jang-Liaw, Chou, Matzke, Iizuka, Min, Kuzmin, Zhang, Cannatella, Hillis, and Che, 2016, Syst. Biol., 65: 835.

English Names

Atlantic Coast Leopard Frog (original publication).

Mid-Atlantic Coast Leopard Frog (Frost, Lemmon, McDiarmid, and Mendelson, 2017, in Crother (ed.), Herpetol. Circ., 43: 15).

Distribution

Central Connecticut and southern New York (where apparently extirpated from Long Island) through New Jersey (presumably adjacent extreme southeastern Pennsylvania) and south along the coastal plain through Delaware, eastern and southwestern Maryland, and tidewater Virginia to North Carolina, USA.

Comment

In the Lithobates pipiens complex, most closely related to Lithobates palustris on the basis of mtDNA evidence according to Newman, Feinberg, Rissler, Burger, and Shaffer, 2012, Mol. Phylogenet. Evol., 63: 445–455, the first discoverers of the unnamed species, although morphologically most similar to Lithobates sphenocephala and Lithobates pipiens. Altig and McDiarmid, 2015, Handb. Larval Amph. US and Canada: 234–235, provided an account of larval morphology and biology (as Lithobates sp.). Schlesinger, Feinberg, Kleopfer, Beane, Bunnell, Burger, Corey, Jaycox, Kiviat, Kubel, Quinn, Raithel, Scott, Wenner, White, Zarate, and Shaffer, 2018, PLoS One, 13(11: e0205805): 1–28, refined the range of the species and it morphological identification.

External links:

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For additional sources of information from other sites search Google
For images search CalPhoto Images and Google Images
To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For related information on conservation and images as well as observation see iNaturalist
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.