Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Lithobates okaloosae (Moler, 1985)

Class: Amphibia > Order: Anura > Family: Ranidae > Genus: Lithobates > Species: Lithobates okaloosae

Rana okaloosae Moler, 1985, Copeia, 1985: 379. Holotype: UF 53964, by original designation. Type locality: "along Malone Creek, Eglin Air Force Base, Okaloosa County, Florida (Sec 24-T2N-R25W), at an altitude of 13 meters".

Rana (Rana) okaloosae — Dubois, 1987 "1986", Alytes, 5: 42, by implication.

Rana (Aquarana) okaloosae — Dubois, 1992, Bull. Mens. Soc. Linn. Lyon, 61: 331; Hillis, 2007, Mol. Phylogenet. Evol., 42: 335-336, by implication and for purposes of discussion; Yuan, Zhou, Chen, Poyarkov, Chen, Jang-Liaw, Chou, Matzke, Iizuka, Min, Kuzmin, Zhang, Cannatella, Hillis, and Che, 2016, Syst. Biol., 65: 835.

Rana (Novirana, Aquarana) okaloosae — Hillis and Wilcox, 2005, Mol. Phylogenet. Evol., 34: 305. See Dubois, 2006, Mol. Phylogenet. Evol., 42: 317-330, Hillis, 2007, Mol. Phylogenet. Evol., 42: 331-338, and Dubois, 2007, Cladistics, 23: 390-402, for relevant discussion of nomenclature. Invalid name formulation under the International Code of Zoological Nomenclature (1999) as discussed by Dubois, 2007, Cladistics, 23: 395.

Rana (Novirana) okaloosae — Hillis and Wilcox, 2005, Mol. Phylogenet. Evol., 34: 305. Nomenclatural act of Hillis and Wilcox, 2005, as interpreted by Fouquette and Dubois, 2014, Checklist N.A. Amph. Rept.: 417.

Lithobates okaloosae — Frost, Grant, Faivovich, Bain, Haas, Haddad, de Sá, Channing, Wilkinson, Donnellan, Raxworthy, Campbell, Blotto, Moler, Drewes, Nussbaum, Lynch, Green, and Wheeler, 2006, Bull. Am. Mus. Nat. Hist., 297: 369; Che, Pang, Zhao, Wu, Zhao, and Zhang, 2007, Mol. Phylogenet. Evol., 43: 1-13; by implication.

Lithobates (Aquarana) okaloosae — Dubois, 2006, C. R. Biol., Paris, 329: 829; Dubois, 2006, Mol. Phylogenet. Evol., 42: 325.

Rana (Lithobates) okaloosae — Fouquette and Dubois, 2014, Checklist N.A. Amph. Rept.: 417.

Aquarana okaloosae — Dubois, Ohler, and Pyron, 2021, Megataxa, 5: 424.

English Names

Florida Bog Frog (Moler, 1985, Copeia, 1985: 379; Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 108; Collins, 1997, Herpetol. Circ., 25: 13; Crother, Boundy, Campbell, de Queiroz, Frost, Highton, Iverson, Meylan, Reeder, Seidel, Sites, Taggart, Tilley, and Wake, 2001 "2000", Herpetol. Circ., 29: 15; Frost, McDiarmid, and Mendelson, 2008, in Crother (ed.), Herpetol. Circ., 37: 8; Collins and Taggart, 2009, Standard Common Curr. Sci. Names N. Am. Amph. Turtles Rept. Crocodil., ed. 6: 8; Frost, McDiarmid, Mendelson, and Green, 2012, in Crother (ed.), Herpetol. Circ., 39: 18; Frost, Lemmon, McDiarmid, and Mendelson, 2017, in Crother (ed.), Herpetol. Circ., 43: 15).

Distribution

Small tributary streams of the East Bay, Shoal, and Yellow Rivers, in Okaloosa, Santa Rosa, and Walton counties, Florida, USA.

Comment

In the Rana catesbeiana group of XXX. In the equivalent Rana clamitans group of Dubois, 1987 "1986", Alytes, 5: 42, and subgenus Aquarana of Dubois, 1992, Bull. Mens. Soc. Linn. Lyon, 61: 331. Reviewed by Moler, 1993, Cat. Am. Amph. Rept., 561: 1-3. Austin, Lougheed, Moler, and Boag, 2003, Biol. J. Linn. Soc., 80, found that population samples of Rana okaloosae are imbedded phylogenetically among populations of Rana clamitans. See photograph, map, description of geographic range and habitat, and conservation status (as Rana okaloosae) in Stuart, Hoffmann, Chanson, Cox, Berridge, Ramani, and Young, 2008, Threatened Amph. World: 509. Dodd, 2013, Frogs U.S. and Canada, 2 : 561–564, provided an account that summarized relevant literature. Elliot, Gerhardt, and Davidson, 2009, Frogs and Toads of N. Am.: 204–205, provided an account, photos, and advertisement call. Altig and McDiarmid, 2015, Handb. Larval Amph. US and Canada: 231–232, provided an account of larval morphology and biology.

External links:

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For related information on conservation and images as well as observation see iNaturalist
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