Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Lithobates pueblae (Zweifel, 1955)

Class: Amphibia > Order: Anura > Family: Ranidae > Genus: Lithobates > Species: Lithobates pueblae

Rana pueblae Zweifel, 1955, Univ. California Publ. Zool., 54: 253. Holotype: UMMZ 99474, by original designation. Type locality: "2. 8 miles northeast of Huauchinango, Río, Texcapa, Puebla, México".

Rana (Rana) pueblae — Dubois, 1987 "1986", Alytes, 5: 42, by implication.

Rana (Zweifelia) pueblae — Dubois, 1992, Bull. Mens. Soc. Linn. Lyon, 61: 331; Hillis, 2007, Mol. Phylogenet. Evol., 42: 335-336, by implication; Yuan, Zhou, Chen, Poyarkov, Chen, Jang-Liaw, Chou, Matzke, Iizuka, Min, Kuzmin, Zhang, Cannatella, Hillis, and Che, 2016, Syst. Biol., 65: 835.

Rana (Novirana, Sierrana, Torrentirana, Zweifelia) pueblae — Hillis and Wilcox, 2005, Mol. Phylogenet. Evol., 34: 305. See Dubois, 2006, Mol. Phylogenet. Evol., 42: 317-330, Hillis, 2007, Mol. Phylogenet. Evol., 42: 331-338, and Dubois, 2007, Cladistics, 23: 390-402, for relevant discussion of nomenclature. Invalid name formulation under the International Code of Zoological Nomenclature (1999) as discussed by Dubois, 2007, Cladistics, 23: 395.

Lithobates pueblae — Frost, Grant, Faivovich, Bain, Haas, Haddad, de Sá, Channing, Wilkinson, Donnellan, Raxworthy, Campbell, Blotto, Moler, Drewes, Nussbaum, Lynch, Green, and Wheeler, 2006, Bull. Am. Mus. Nat. Hist., 297: 369. Che, Pang, Zhao, Wu, Zhao, and Zhang, 2007, Mol. Phylogenet. Evol., 43: 1-13; by implication.

Lithobates (Lithobates) pueblae — Dubois, 2006, C. R. Biol., Paris, 329: 830; Dubois, 2006, Mol. Phylogenet. Evol., 42: 325.

Common Names

Puebla Frog (Liner, 1994, Herpetol. Circ., 23: 27; Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 109; Liner and Casas-Andreu, 2008, Herpetol. Circ., 38: 16).

Distribution

Known only from the vicinity of Huauchinango and adjacent Río Nexcapa in northern Puebla, Mexico.

Geographic Occurrence

Natural Resident: Mexico

Endemic: Mexico

Comment

In the Rana tarahumarae group of Hillis, Frost, and Webb, 1984, Copeia, 1984: 398-403, Rana pustulosa group of Dubois, 1987 "1986", Alytes, 5: 42. In the subgenus Zweifelia of Dubois, 1992, Bull. Mens. Soc. Linn. Lyon, 61: 331. See account by Webb, 1988, Occas. Pap. Mus. Texas Tech Univ., 121: 1-15. See map, description of geographic range and habitat, and conservation status (as Rana pueblae) in Stuart, Hoffmann, Chanson, Cox, Berridge, Ramani, and Young, 2008, Threatened Amph. World: 510.

External links:

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For access to general information see Wikipedia
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For images search CalPhoto Images and Google Images
To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.