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Rana tsushimensis Stejneger, 1907
Rana tsushimensis Stejneger, 1907, Bull. U.S. Natl. Mus., 58: 116. Holotype: USNM 17519 by original designation and Cochran, 1961, Bull. U.S. Natl. Mus., 220: 77. Type locality: "Tsushima, in the Straits of Korea, between the south end of Korea and Kiusiu, Japan".
Rana temporaria tsushimensis — Sato and Takashima, 1955, Misc. Rep. Yamashina Inst. Ornithol. Zool., 6: 262; Kawamura, 1962, J. Sci. Hiroshima Univ., B—Zool., 20: 181-193.
Rana tsushimensis — Kawamura, 1962, J. Sci. Hiroshima Univ., B—Zool., 20: 191; Okada, 1966, Fauna Japon., Anura: 95; Nishioka, Okumoto, Ueda, and Ryuzaki, 1987, Sci. Rep. Lab. Amph. Biol. Hiroshima Univ., 9: 165-212.
Rana (Rana) amurensis tsushimensis — Nakamura and Ueno, 1963, Japan. Rept. Amph. Color: 37.
Rana amurensis tsushimensis — Ueno and Shibata, 1970, Mem. Nat. Sci. Mus. Tokyo, 3: 194.
Rana (Rana) tsushimensis — Dubois, 1987 "1986", Alytes, 5: 41, by implication; Dubois, 1992, Bull. Mens. Soc. Linn. Lyon, 61: 333; Yuan, Zhou, Chen, Poyarkov, Chen, Jang-Liaw, Chou, Matzke, Iizuka, Min, Kuzmin, Zhang, Cannatella, Hillis, and Che, 2016, Syst. Biol., 65: 835.
Rana (Laurasiarana) tsishimensis — Hillis and Wilcox, 2005, Mol. Phylogenet. Evol., 34: 311, by implication; see Dubois, 2006, Mol. Phylogenet. Evol., 42: 317-330, and Hillis, 2007, Mol. Phylogenet. Evol., 42: 331-338, for discussion.
Common Names
Tsushima Leopard Frog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 110).
Tsushima Brown Frog (Maeda and Matsui, 1990, Frogs Toads Japan, Ed. 2: 51; Goris and Maeda, 2004, Guide Amph. Rept. Japan: 105).
Distribution
Tsushima I., in the Korea Strait, Japan.
Geographic Occurrence
Natural Resident: Japan
Endemic: Japan
Comment
In the Section Rana, subgenus Rana, Rana temporaria group of Dubois, 1992, Bull. Mens. Soc. Linn. Lyon, 61: 333. See Kuramoto, 1974, Copeia, 1974: 815-822, for a discussion of relationships. Reviewed by Okada, 1966, Fauna Japon., Anura: 95-99. Removed from the synonymy of Rana japonica by Kawamura, 1962, J. Sci. Hiroshima Univ., B—Zool., 20: 191, where it had been placed by Boulenger, 1920, Rec. Indian Mus., 20: 93. See Maeda and Matsui, 1989, Frogs Toads Japan, Ed. 1: 51. Most closely related to Rana okinavana according to Tanaka, Matsui, and Takenaka, 1996, Biochem. Syst. Ecol., 24: 302. Maeda and Matsui, 1990, Frogs Toads Japan, Ed. 2: 48-51, provided an account. Goris and Maeda, 2004, Guide Amph. Rept. Japan: 105-107, provided an account, map, and photograph.
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