Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Philautus macroscelis (Boulenger, 1896)

Class: Amphibia > Order: Anura > Family: Rhacophoridae > Subfamily: Rhacophorinae > Genus: Philautus > Species: Philautus macroscelis

Rhacophorus macroscelis Boulenger, 1896, Ann. Mag. Nat. Hist., Ser. 6, 17: 403. Holotype: BMNH, by original designation. Holotype is BMNH 1947.2.8.85 (formerly numbered 1895.11.7.82) by museum records. Type locality: "Mount Kina Balu, N. Borneo [= Sabah]".

Philautus spiculatus Smith, 1931, Bull. Raffles Mus., 5: 20. Holotype: BMNH, by original designation, by museum records this is BMNH 1947.2.6.4 (originally 1929.12.22.24). Type locality: "Kenokok, alt. 3,300 feet", Mount Kina Balu, Sabah, Malaysia (Borneo). Synonymy with Rhacophorus everetti (sensu lato) by Inger, 1966, Fieldiana, Zool., 52: 298–300. Synonymy with Rhacophorus macroscelis implied by Hertwig, Das, Schweizer, Brown, and Haas, 2012, Zool. Scripta, 41: 29–46.

Rhacophorus (Rhacophorus) everetti — Ahl, 1931, Das Tierreich, 55: 119.

Rhacophorus (Rhacophorus) macroscelis — Ahl, 1931, Das Tierreich, 55: 119.

Philautus spiculosus — Smith, 1931, Bull. Raffles Mus., 5: 32. Incorrect subsequent spelling.

Rhacophorus spiculatus — Inger, 1954, J. Washington Acad. Sci., 44: 251.

Rhacophorus everetti macroscelis — Inger, 1954, Fieldiana, Zool., 33: 385.

Philautus macroscelis — Hertwig, Das, Schweizer, Brown, and Haas, 2012, Zool. Scripta, 41: 29.

English Names

Mossy Bush Frog (Haas, Das, Hertwig, Bublies, and Schulz-Schaeffer, 2022, Guide to the Tadpoles of Borneo: 364).

Distribution

Borneo in the mountainous area from Gunung Kinabalu National Park in Sabah to Gunung Mulu National Park in Sarawak, Malaysia, and Brunei, 750–1800 m elevation; possibly to be found in adjacent Kalimantan, Indonesia.

Comment

Removed from the synonymy of Rhacophorus everetti by Hertwig, Das, Schweizer, Brown, and Haas, 2012, Zool. Scripta, 41: 29–46, where it had been placed by Inger, 1966, Fieldiana, Zool., 52: 297. Formerly placed as a synonym of Rhacophorus buergeri hosii by Wolf, 1936, Bull. Raffles Mus., 12: 170. See account for Bornean population (as Rhacophorus everetti macroscelis) by Inger, 1966, Fieldiana, Zool., 52: 298–300. Malkmus, Manthey, Vogel, Hoffmann, and Kosuch, 2002, Amph. Rept. Mount Kinabalu: 210–212, provided an account as Rhacophorus everetti macroscelis. Manthey and Grossmann, 1997, Amph. Rept. Südostasiens: 122–123, to compare this species to other rhacophorids of the Sunda Shelf region (in the sense of including Rhacophorus macroscelis). Haas, Das, Hertwig, Bublies, and Schulz-Schaeffer, 2022, Guide to the Tadpoles of Borneo: 364–366, summarized the knowledge of habitat, reproduction, larval morphology and coloration.

External links:

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For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For related information on conservation and images as well as observation see iNaturalist; for a quick link to their maps see iNaturalist KML
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.