Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Philautus worcesteri (Stejneger, 1905)

Class: Amphibia > Order: Anura > Family: Rhacophoridae > Subfamily: Rhacophorinae > Genus: Philautus > Species: Philautus worcesteri

Cornufer worcesteri Stejneger, 1905, Proc. U.S. Natl. Mus., 28: 345. Holotype: USNM 34784 by original designation. Type locality: "Mount Apo, 6,0000 feet altitude, Mindanao [Island], Philippine Islands".

Rhacophorus emembranatus Inger, 1954, Fieldiana, Zool., 33: 392. Holotype: FMNH 50684, by original designation. Type locality: "east slope of Mount McKinley, Mindanao, altitude 950 meters", Philippines. Synonymy by Brown, Alcala, and Brown, 1998, J. Herpetol., 32: 131–132.

Philautus emembranatus — Liem, 1970, Fieldiana, Zool., 57: 68; Inger, 1985, in Frost (ed.), Amph. Species World: 528.

Philautus (Philautus) emembranatus — Dubois, 1987 "1986", Alytes, 5: 72.

Philautus worcesteri — Brown, Alcala, and Brown, 1998, J. Herpetol., 32: 131.

English Names

Mindanao Bubble-nest Frog (Philautus emembranatus [no longer recognized]: Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 112).

Distribution

Mountain ranges in Agusan del Norte, Davao, Misamis Occidental, and Zamboanga del Norte provinces, Mindanao, Philippines.

Geographic Occurrence

Natural Resident: Philippines

Endemic: Philippines

Comment

See account by Brown and Alcala, 1994, Proc. California Acad. Sci., Ser. 4, 48: 185–220 (as Philautus emembranatus, and who considered it to be in the Philautus surdus group). Brown, Alcala, and Brown, 1998, J. Herpetol., 32: 131–132, discussed the nomenclature of the species, removing the senior name from the synonymy of Platymantis guentheri, where it had been placed by Inger, 1954, Fieldiana, Zool., 33: 362. See photograph, map, description of geographic range and habitat, and conservation status in Stuart, Hoffmann, Chanson, Cox, Berridge, Ramani, and Young, 2008, Threatened Amph. World: 535. See brief notes for Mindanao, Philippines, by Sanguila, Cobb, Siler, Diesmos, Alcala, and Brown, 2016, ZooKeys, 624: 49–50. See comments by Baron, Marin, Logramonte, and Mohagan, 2021, Asian Herpetol. Res., 12: 76–87.

External links:

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For additional sources of general information from other websites search Google
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For images search CalPhoto Images and Google Images
To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observation see iNaturalist
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.