Zhangixalus dulitensis (Boulenger, 1892)

Class: Amphibia > Order: Anura > Family: Rhacophoridae > Subfamily: Rhacophorinae > Genus: Zhangixalus > Species: Zhangixalus dulitensis

Rhacophorus dulitensis Boulenger, 1892, Proc. Zool. Soc. London, 1892: 507. Holotype: BMNH 1892.6.3.15, now renumbered as 1947.2.8.84 according to museum records. Type locality: "Mt. Dulit, Sarawak", Malaysia (Borneo).

Rhacophorus chiropterus Werner, 1896, Verh. Zool. Bot. Ges. Wien, 46: 18. Holotype: ZMB. Type locality: "Nord Borneo", Malaysia. Synonymy by Boulenger, 1897, Zool. Rec., 33: 34; Van Kampen, 1923, Amph. Indo-Austral. Arch.: 258; Ahl, 1931, Das Tierreich, 55: 173; Wolf, 1936, Bull. Raffles Mus., 12: 210.

Rhacophorus (Rhacophorus) dulitensisAhl, 1931, Das Tierreich, 55: 173; Dubois, 1987 "1986", Alytes, 5: 77.

Zhangixalus dulitensis — Jiang, Jiang, Ren, Wu, and Li, 2019, Asian Herpetol. Res., 10: 7. 

Rhacophorus (Zhangixalus) dulitensis — Mahony, Kamei, Brown, and Chan, 2024, Vert. Zool., Senckenberg, 74: 253, by implication. 

English Names

Dulit Flying Frog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 113).

Jade Tree Frog (Das, 2007, Amph. Rept. Brunei: 76).


Isolated populations in the northern half of Borneo (Sabah and Sarawak, Malaysia, Brunei, and East, Central, and West Kalimantan provinces, Indonesia), below 250 m elevation.

Geographic Occurrence

Natural Resident: Brunei, Indonesia, Malaysia, Malaysia, East (Sarawak and/or Sabah)


In the Rhacophorus (Rhacophorus) reinwardtii group of Dubois, 1987 "1986", Alytes, 5: 77; see comment under Rhacophorus for dissenting opinion regarding the recognition of this group. See account by Bourret, 1942, Batr. Indochine: 448–449. See Inger, 1966, Fieldiana, Zool., 52: 293–297, for account. See identification table by Manthey and Grossmann, 1997, Amph. Rept. Südostasiens: 122–123, to compare this species to other rhacophorids of the Sunda Shelf region. Malkmus, Manthey, Vogel, Hoffmann, and Kosuch, 2002, Amph. Rept. Mount Kinabalu: 209–210, provided an account. See statement of geographic range, habitat, and conservation status in Stuart, Hoffmann, Chanson, Cox, Berridge, Ramani, and Young, 2008, Threatened Amph. World: 635, who noted that the records for Rhacophorus dulitensis for Sumatra are now referred to Rhacophorus prominanus. Das, 2007, Amph. Rept. Brunei: 76, provided a photograph and brief account. Haas, Hertwig, Krings, Braskamp, Dehling, Min, Jankowski, Schweizer, and Das, 2012, Zootaxa, 3328: 1–19, reported on larval morphology. Dehling and Dehling, 2013, Check List, 9: 425–426, provided a range extension to western Borneo and provided a range map. Ujang, Hadimar, Llano-Sánchez, Maiwald, and Das, 2021, Herpetol. Rev., 52: 79, provided a record from West Kalimantan Province (Borneo), Indonesia. Gillespie, Ahmad, and Shia, 2021, Field Guide Frog Lower Kinabatangan Region Sabah: 41, provided a brief account, summarizing identification and life history in the Lower Kinabatangan Region, Sabah, Malaysia. Haas, Das, Hertwig, Bublies, and Schulz-Schaeffer, 2022, Guide to the Tadpoles of Borneo: 394–396, summarized the knowledge of habitat, reproduction, larval morphology and coloration. 

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