Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Zhangixalus melanoleucus Brakels, Nguyen, Pawangkhanant, Idiiatullina, Lorphengsy, Suwannapoom, and Poyarkov, 2023

Class: Amphibia > Order: Anura > Family: Rhacophoridae > Subfamily: Rhacophorinae > Genus: Zhangixalus > Species: Zhangixalus melanoleucus

Zhangixalus melanoleucus Brakels, Nguyen, Pawangkhanant, Idiiatullina, Lorphengsy, Suwannapoom, and Poyarkov, 2023, Zool. Res., Kunming, 44: 376. Holotype: BEI 01010, by original designation. Type locality: "from the montane evergreen forest on Phou Samsoum Mountain, Xiengkhouang Province, northeast Laos (N19.13101°, E103.78408°; at an elevation of 2 066 m a.s.l.)". Zoobank publication registrations: 601DF810-CCDE-4D0A-A8AF-81837DDE02EA

Rhacophorus (Zhangixalus) melanoleucus — Mahony, Kamei, Brown, and Chan, 2024, Vert. Zool., Senckenberg, 74: 253.

Common Names

Phou Samsoum Treefrog (original publication).

Distribution

Known only from the type locality in high-elevation montane evergreen forest of Phou Samsoum Mountain, Xiengkhouang Province, Laos; expected to be found in other mountain areas in the Houaphan, Xaisomboun, and Bolikhamxay provinces of Laos and adjacent Nghe An Province in Vietnam.

Geographic Occurrence

Natural Resident: Laos

Likely/Controversially Present: Vietnam

Endemic: Laos

Comment

The sister species of Zhangixalus nigropunctatus, according to the original publication, where comparative morphology, molecular markers, acoustics, and natural history were detailed. Stuart, Seateun, Sivongxay, and Phimmachak, 2025 "2024", in Wildlife Lao PDR: 45, summarized (as Rhacophorus melanoleucus) literature, taxonomy, habitat, and detailed range for Laos.

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