Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Bolitoglossa biseriata Tanner, 1962

Class: Amphibia > Order: Caudata > Family: Plethodontidae > Subfamily: Hemidactyliinae > Genus: Bolitoglossa > Species: Bolitoglossa biseriata

Bolitoglossa biseriata Tanner, 1962, Herpetologica, 18: 19. Holotype: UU 3847, by original designation. Type locality: "Rio Tuira at mouth of Paya Tributary, Darien Province, Panama".

Bolitoglossa (Eladinea) biseriata — Parra-Olea, García-París, and Wake, 2004, Biol. J. Linn. Soc., 81: 336.

Common Names

Two-lined Mushroomtongue Salamander (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 29).

Distribution

Northwestern, south-central, and eastern Panama (Provinces of Bocas del Toro, Coclé, Panamá, San Blas [now Kuna Yala Indian Territory], and Darién) and western Colombia (Departments of Valle, Cauca, and Nariño) south, possibly, to northwestern Ecuador (see comment).

Geographic Occurrence

Natural Resident: Colombia, Panama

Likely/Controversially Present: Ecuador

Comment

In the Bolitoglossa (Eladinea) adspersa group of Parra-Olea, García-París, and Wake, 2004, Biol. J. Linn. Soc., 81: 336; formerly in the Bolitoglossa sima group of Wake and Lynch, 1976, Sci. Bull. Nat. Hist. Mus. Los Angeles Co., 25: 1-65. See account by Brame and Wake, 1963, Contrib. Sci. Nat. Hist. Mus. Los Angeles Co., 69: 38. Cisneros-Heredia, 2007, Check List, 2(3): 64-67, provided a discussion of the range as well as a record for Ecuador. Mueses-Cisneros and Moreno-Quintero, 2012, Herpetotropicos, Mérida, 7: 39-54, provided the first record for Nariño, southwestern Colombia. Köhler, 2011, Amph. Cent. Am.: 40–69, compared this species with others from Central America, provided a map, photograph, and brief characterization. Raffaëlli, 2013, Urodeles du Monde, 2nd ed.: 340, provided a brief account, photograph, and range map. Raffaëlli, 2022, Salamanders & Newts of the World: 739–740, provided an account summarizing systematics, life history, population status, and distribution (including a polygon map). Coloma and Duellman, 2025, Amph. Ecuador. Vol. 4: 365, did not include this species as a member of the Ecuadorian fauna noting ambiguity in the evidence published so far with some (e.g., Hervas, 2016, Unpubl. Dissertation) suggesting that the Ecuador population is assignable to Bolitoglossa sima.

External links:

Please note: these links will take you to external websites not affiliated with the American Museum of Natural History. We are not responsible for their content.

For access to general information see Wikipedia
For additional sources of general information from other websites search Google
For access to relevant technical literature search Google Scholar
For images search CalPhoto Images and Google Images
To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For related information on conservation and images as well as observations see iNaturalist
For additional information specific to Ecuador see FaunaWebEcuador: Anfibios del Ecuador