Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Bolitoglossa gracilis Bolaños, Robinson, and Wake, 1987

Class: Amphibia > Order: Caudata > Family: Plethodontidae > Subfamily: Hemidactyliinae > Genus: Bolitoglossa > Species: Bolitoglossa gracilis

Bolitoglossa gracilis Bolaños, Robinson, and Wake, 1987, Rev. Biol. Tropical, 35: 87. Holotype: UCR 9378, by original designation. Type locality: "Río Quirí, about 1 km NE by road from the bridge crossing the Río Grande de Orosi near Tapantí, Cartago Province, Costa Rica, at an elevation of approximately 1, 280 m . . . 9° 47′ 30″ N, 83° 47′ 42″ W, 1280 m".

Bolitoglossa (Eladinea) gracilis — Parra-Olea, García-París, and Wake, 2004, Biol. J. Linn. Soc., 81: 335.

Common Names

Rio Quiri Salamander (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 29).

Rio Quiri Mushroomtongue Salamander (Raffaëlli, 2022, Salamanders & Newts of the World: 683).

Distribution

Vicinity of the type locality near Tapantí in lower montane rainforest (1225–1280 m elevation) on the Atlantic slope of Cartago Province, central Costa Rica; also reported from two sites on the eastern slope of the Volcán Turrialba, Rio Vereh Cloud Forest Reserve, and the Caribbean slopes of the Sierra de Talamanca at 1400–1700 m elevation, suggesting that the species may enter Panama.

Geographic Occurrence

Natural Resident: Costa Rica

Likely/Controversially Present: Panama

Endemic: Costa Rica

Comment

In the Bolitoglossa subpalmata group by implication. Most similar to Bolitoglossa diminuta and Bolitoglossa subpalmata according to the original publication. See account by Savage, 2002, Amph. Rept. Costa Rica: 133. In the Bolitoglossa (Eladinea) subpalmata group of Parra-Olea, García-París, and Wake, 2004, Biol. J. Linn. Soc., 81: 336. See García-París, Parra-Olea, and Wake, 2008, Herpetol. J., 18: 23-31, for comments on systematics. See photograph, map, description of geographic range and habitat, and conservation status in Stuart, Hoffmann, Chanson, Cox, Berridge, Ramani, and Young, 2008, Threatened Amph. World: 558. Köhler, 2011, Amph. Cent. Am.: 40–69, compared this species with others from Central America and provided a map and photograph. Raffaëlli, 2013, Urodeles du Monde, 2nd ed.: 320, provided a brief account, photograph, and map. Raffaëlli, 2022, Salamanders & Newts of the World: 683, provided an account summarizing systematics, life history, population status, and distribution (including a polygon map), as well as noting likely unvouchered records from Volcán Turrialba, Río Vereh, and the Sierra Talamanca, eastern Costa Rica.

External links:

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For access to general information see Wikipedia
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For images search CalPhoto Images and Google Images
To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist