Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Necturus lewisi Brimley, 1924

Class: Amphibia > Order: Caudata > Family: Proteidae > Genus: Necturus > Species: Necturus lewisi

Necturus maculosus lewisi Brimley, 1924, J. Elisha Mitchell Sci. Soc., 40: 167. Holotype: Brimley 6868, by original designation; now USNM 73848, according to Tighe, 2022, Smithson. Contrib. Zool., 654: 66. Type locality: "Neuse River, near Raleigh", Wake County, North Carolina, USA.

Necturus lewisi — Bishop, 1943, Handb. Salamanders: 32; Hecht, 1958, Proc. Staten Island Inst. Arts Sci., 21: 15.

Necturus maculosus lewisi — Schmidt, 1953, Check List N. Am. Amph. Rept., Ed. 6: 13; Brode, 1970, Dissert. Abstr. Internatl., Ser. B, 30: 5288-5289.

Necturus (Necturus) lewisi — Dubois and Raffaëlli, 2012, Alytes, 28: 147.

Common Names

Lewis' Mudpuppy (Bishop, 1943, Handb. Salamanders: 32).

Neuse River Waterdog (Schmidt, 1953, Check List N. Am. Amph. Rept., Ed. 6: 13; Conant, Cagle, Goin, Lowe, Neill, Netting, Schmidt, Shaw, Stebbins, and Bogert, 1956, Copeia, 1956: 175; Conant, 1975, Field Guide Rept. Amph. E. Cent. N. Am., Ed. 2: 244; Collins, Huheey, Knight, and Smith, 1978, Herpetol. Circ., 7: 7; Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 34; Collins, 1997, Herpetol. Circ., 25: 8; Crother, Boundy, Campbell, de Queiroz, Frost, Highton, Iverson, Meylan, Reeder, Seidel, Sites, Taggart, Tilley, and Wake, 2001 "2000", Herpetol. Circ., 29: 25; Collins and Taggart, 2009, Standard Common Curr. Sci. Names N. Am. Amph. Turtles Rept. Crocodil., ed. 6: 15; Tilley, Highton, and Wake, 2012, in Crother (ed.), Herpetol. Circ., 39: 28; Highton, Bonett, and Jockusch, 2017, in Crother (ed.), Herpetol. Circ., 43: 30).

Distribution

Neuse and Tar-Pamlico River systems, North Carolina, USA, sea level–160 m elevation. See comment.

Geographic Occurrence

Natural Resident: United States of America, United States of America - North Carolina

Endemic: United States of America, United States of America - North Carolina

Comment

See accounts by Ashton, 1990, Cat. Am. Amph. Rept., 456: 1–2, and Petranka, 1998, Salamand. U.S. Canada: 422–425. See also Ashton, Braswell, and Guttman, 1981 "1980", Brimleyana, 4: 43–46, for evidence in support of the distinctiveness of this taxon from Necturus maculosus and Necturus punctatus. Brode, 1970, Dissert. Abstr. Internatl., Ser. B, 30: 5288–5289, suggested that Necturus lewisi (as Necturus maculosus lewisi) and Necturus maculosus (as Necturus maculosus maculosus) formed an intergrading series of populations. See also Guttman, Weigt, Moler, Ashton, Mansell, and Peavy, 1990, J. Herpetol., 24: 163–175, for geographic variation in allozymes. Braswell, 2005, in Lannoo (ed.), Amph. Declines: 868–870, provided a detailed account that summarized the biology and conservation literature. See statement of geographic range, habitat, and conservation status in Stuart, Hoffmann, Chanson, Cox, Berridge, Ramani, and Young, 2008, Threatened Amph. World: 641. Raffaëlli, 2013, Urodeles du Monde, 2nd ed.: 217–218, provided a brief account, photograph, and map. Altig and McDiarmid, 2015, Handb. Larval Amph. US and Canada: 127–128, provided an account of larval morphology and biology. Nelson, Niemiller, and Fitzpatrick, 2017, J. Herpetol., 51: 559–566, reported a population of Necturus aff. lewisi from the western side of the Appalachian range in eastern Tennessee, which seemingly is involved in ongoing hybridization with Necturus maculosus. Raffaëlli, 2022, Salamanders & Newts of the World: 476–478, provided an account, summarizing systematics, life history, population status, and distribution (including a polygon map) as well as a brief discussion of Necturus aff. lewisi from Tennessee, USA. Tighe, 2022, Smithson. Contrib. Zool., 654: 66, briefly discussed location of paratypes.

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To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist