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* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
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Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Triturus macedonicus (Karaman, 1922)

Class: Amphibia > Order: Caudata > Family: Salamandridae > Subfamily: Pleurodelinae > Genus: Triturus > Species: Triturus macedonicus

Molge karelinii var. macedonica Karaman, 1922, Glasn. Naucni Casopis za Prirodne Nauke, Zagreb, 34: 280. Syntypes: Deposition not stated. Type locality: "Ohrid", Republic of Macedonia.

Triturus carnifex macedonicus — Arntzen and Wallis, 1999, Bijdr. Dierkd., 68: 181–203, by implication.

Triturus macedonicus — Arntzen, Espregueira Themudo, and Wielstra, 2007, Contrib. Zool., Amsterdam, 76: 275.

Triturus (Triturus) macedonicus — Dubois and Raffaëlli, 2009, Alytes, 26: 67.

Triturus cristatus carnifex var. albanicus Dely, 1959, Ann. Hist. Nat. Mus. Natl. Hungarici, 51: 445. Holotype: MNH 58.819.1 (no: cr 4), by original designation. Type locality: "Mali me Grope Stanet e Shen Merise (Albania)". Unavailable tetranominal.

Triturus karelinii arntzeni Litvinchuk, Borkin, Džukić and Kalezić in Litvinchuk, Borkin, Džukić, Kalezić, Khalturin, and Rosanov, 1999, Russ. J. Herpetol., 6: 159. Holotype: ZISP 6121, by original designation. Type locality: "vicinity of the village Vrtovac (43° 25´ N, 22° 28´ E, UTM FP10), altitude 400 m above sea level", Serbia. Synonymy by Wielstra, Litvinchuk, Naumov, Tzankov, and Arntzen, 2013, Zootaxa, 3682 : 443. Consideration as a hybrid population of Triturus macedonicus X Triturus ivanbureschi by Wielstra and Arntzen, 2014, Zootaxa, 3802: 381.

Triturus (Triturus) karelinii arntzeni — Dubois and Raffaëlli, 2009, Alytes, 26: 67. See comment above.

Triturus arntzeni — Espregueira Themudo, Wielstra, and Arntzen, 2009, Mol. Phylogenet. Evol., 52: 328. See comment above.

Trturus (Triturus) macedonicus — Raffaëlli, 2022, Salamanders & Newts of the World: 376.

English Names

Macedonian Crested Newt (Speybroeck, Beukema, Bok, and Van Der Voort, 2016, Field Guide Amph. Rept. Brit. Eur.: 97).

Distribution

Eastern Bosnia-Herzegovina, central and southern Serbia, Montenegro, central and western North Macedonia and Albania to northeastern Bulgaria, south into northwestern and central Greece.

Comment

Arntzen, Espregueira Themudo, and Wielstra, 2007, Contrib. Zool., Amsterdam, 76: 261–278, distinguished Triturus arntzeni from Triturus carnifex and provided an account. Triturus arntzeni was reviewed by Jehle, Thiesmeier, and Foster, 2011, The Crested Newt.: 1–152. Stojanov, Tzankov, and Naumov, 2011, Die Amph. Rept. Bulgariens: 126–130, provided a fairly detailed account of Triturus arntzeni and range map for Bulgaria. Wielstra and Arntzen, 2011, BMC Evol. Biol., 11(162): 1–8, reported ln the molecular phylogenetics of the species and its near relatives. Wielstra and Arntzen, 2012, BMC Evol. Biol., 12(161): 1–12, discussed the mtDNA introgression between Triturus macedonicus and Triturus karelinii. Espregueira Themudo, Wielstra, and Arntzen, 2009, Mol. Phylogenet. Evol., 52: 321–328, noted that the contact zone with Triturus karelini is not well-understood and may be in European Turkey or in western Anatolia. See comments under Triturus karelini. Valakos, Pafilis, Sotiropoulos, Lymberakis, Maragou, and Foufopoulos, 2008, Amph. Rept. Greece: 159–162, provided an account for Greece (as Triturus carnifex). Wielstra, Espregueira Themudo, Güçlü, Olgun, Poyarkov, and Arntzen, 2010, Mol. Phylogenet. Evol., 56: 888–896, reported on the phylogeography of this species along with its sister taxon, Triturus karelinii, and supported its distinctiveness. Arntzen and Wielstra, 2010, Amphibia-Reptilia, 31: 311–322, discussed the taxonomic recognition of this species. Reviewed by Jehle, Thiesmeier, and Foster, 2011, The Crested Newt.: 1–152. Stojanov, Tzankov, and Naumov, 2011, Die Amph. Rept. Bulgariens: 110–119, provided a fairly detailed account (as Triturus karelini) and range map for Bulgaria. Wielstra and Arntzen, 2014, Zootaxa, 3802: 381–388, considered Triturus arntzeni as a hybrid population of Triturus macedonicus X Triturus ivanbureschi. Raffaëlli, 2013, Urodeles du Monde, 2nd ed.: 132, provided a brief account, photo, and map. Wielstra, Sillero, Vörös, and Arntzen, 2014, Amphibia-Reptilia, 35: 376–381, provided a dot map. Arntzen, Wielstra, and Wallis, 2014, Biol. J. Linn. Soc., 113: 604–622, reported on hybrid/contact zones with Triturus cristatus, Triturus carnifex, Triturus dobrogicus, and Triturus ivanbureschi. Speybroeck, Beukema, Bok, and Van Der Voort, 2016, Field Guide Amph. Rept. Brit. Eur.: 97–101, provided a brief account and distribution map. Szabolcs, Mizsei, Jablonski, Vági, Mester, Végvári, and Lengyel, 2017, Amphibia-Reptilia, 38: 435–448, provided a dot map and discussion of range in Albania. See Dufresnes, 2019, Amph. Eur., N. Afr., & Middle East: 136, for brief summary of identifying morphology and biology, a range map, as well as a photograph. Gvozdenović and Iković, 2022, Nat. Montenegrina, Podgorica, 14: 95, included this species in the Montenegro fauna. Raffaëlli, 2022, Salamanders & Newts of the World: 376–377, provided an account, summarizing systematics, life history, population status, and distribution (including a polygon map).

External links:

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For additional sources of information from other sites search Google
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To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For related information on conservation and images as well as observation see iNaturalist
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.