Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Tylototriton anguliceps Le, Nguyen, Nishikawa, Nguyen, Pham, Matsui, Bernardes, and Nguyen, 2015

Class: Amphibia > Order: Caudata > Family: Salamandridae > Subfamily: Pleurodelinae > Genus: Tylototriton > Species: Tylototriton anguliceps

Tylototriton anguliceps Le, Nguyen, Nishikawa, Nguyen, Pham, Matsui, Bernardes, and Nguyen, 2015, Curr. Herpetol., Kyoto, 34: 45. Holotype: HNUE A.I.1.109, by original designation. Type locality: "Muong Nhe Nature Reserve, Muong Nhe District, Dien Bien Province, Vietnam (22° 18.580′ N, 102° 11.026′ E, elevation 1704 m asl)".

Tylototriton (Tylototriton) anguliceps — Raffaëlli, 2022, Salamanders & Newts of the World: 312.

Common Names

Angular-headed Newt (original publication).

Angle-headed Crocodile Newt (Stuart, Seateun, Sivongxay, and Phimmachak, 2025 "2024", in Wildlife Lao PDR: 20).

Angular-headed Corcodile [sic] Newt (Poyarkov, Nguyen, Popov, Geissler, Pawangkhanant, Neang, Suwannapoom, and Orlov, 2021, Russ. J. Herpetol., 28 (3A): 66).

Muong Nhe Crocodile Newt (Raffaëlli, 2022, Salamanders & Newts of the World: 312).

Distribution

Chiang Mai, Chiang Rai, and Nan provinces, Thailand; Phou Ya Kaho, Viengphoukha District, Luang Namtha Province, Laos; and Muong Nhe, Dien Bien Province, and Thuan Chau, Son La Province, northern Vietnam; presumably in adjacent southwestern Yunnan, China, and easternmost Myanmar.

Geographic Occurrence

Natural Resident: Laos, Thailand, Vietnam

Likely/Controversially Present: China, People's Republic of, Myanmar

Comment

Nested within the clade comprised of Tylototriton uyenoi, Tylototriton shanjing, Tylototriton verrucosus, and Tylototriton yangi, according to the original publication. See brief account by Phimmachak, Aowphol, and Stuart, 2015, Zootaxa, 4006: 297–299, who provided the Laos locality. Hernandez, Escoriza, Pomchote, and Hou, 2019, Herpetol. Bull., London, 147: 15–18, provided a record from Chiang Mai province, Thailand, and discussed the habitat and range. Niyomwan, Srisom, and Pawangkhanant, 2019, Field Guide Amph. Thailand: 92–93, provided a brief account (photograph, habitat, and range) for Thailand (in Thai). Bernardes, Le, Nguyen, Pham, Pham, Nguyen, Rödder, Bonkowski, and Ziegler, 2020, ZooKeys, 935: 121–164, provided a range map. Makchai, Chuaynkern, Safoowong, Chuachat, and Cota, 2020, Amph. N. Thailand: 185, provided photographs, a brief account for Thailand, and a range map. Makchai, Chuaynkern, Safoowong, Chuachat, and Cota, 2020, Amph. N. Thailand: 185, provided photographs, a brief account for Thailand, and a range map. Hernandez and Pomchote, 2020, Herpetol. Notes, 13: 993–996, reported the species from Khun Chae National Park, Chiang Rai Province, Thailand, and discussed the range in Thailand. In the Tylototriton (Tylototriton) verrucosus species group of Poyarkov, Nguyen, and Arkhipov, 2021, Taprobanica, 10: 4–22, who discussed phylogenetics. In the Tylototriton verrucosus group of Lyu, Wang, Zeng, Zhou, Qi, Wan, Li, and Wang, 2021, Vert. Zool., Senckenberg, 71: 697–710, who discussed phylogenetics. Raffaëlli, 2022, Salamanders & Newts of the World: 312–313, provided an account, summarizing systematics, life history, population status, and distribution (including a polygon map). Pham, Nguyen, Nguyen, Le, Ziegler, and Tran, 2024, Amph. Rept. Conserv., 18: 20–47, discussed natural history and records from Copia Nature Reserve, Son La Province, Vietnam. Stuart, Seateun, Sivongxay, and Phimmachak, 2025 "2024", in Wildlife Lao PDR: 20, briefly summarized literature, taxonomy, habitat, and range for Laos.

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