Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Tylototriton notialis Stuart, Phimmachak, Sivongxay, and Robichaud, 2010

Class: Amphibia > Order: Caudata > Family: Salamandridae > Subfamily: Pleurodelinae > Genus: Tylototriton > Species: Tylototriton notialis

Tylototriton (Yaotriton) notialis Stuart, Phimmachak, Sivongxay, and Robichaud, 2010, Zootaxa, 2650: 23. Holotype: FMNH 271121, by original designation. Type locality: "Laos, Khammouan Province, Boualapha District, Nakai-Nam Theun National Protected Area, Nam On River catchment, Phou Ak escarpment, 17° 38′ 39.6″ N 105° 44′ 12.3″ E (Fig. 4), 980 m elev."

Yaotriton notialis — Dubois, Ohler, and Pyron, 2021, Megataxa, 5: 408.

Common Names

Southern Crocodile Newt (Stuart, Seateun, Sivongxay, and Phimmachak, 2025 "2024", in Wildlife Lao PDR: 20).

Southern Corcodile [sic] Newt (Poyarkov, Nguyen, Popov, Geissler, Pawangkhanant, Neang, Suwannapoom, and Orlov, 2021, Russ. J. Herpetol., 28 (3A): 67).

Khammouan Crocodile Newt (Raffaëlli, 2022, Salamanders & Newts of the World: 288).

Laos Knobby Newt (Sparreboom, 2014, Salamanders Old World: 365).

Distribution

Three known localities in the Northern Annamites, one in Xaichamphone District, Bolikhamxai Province, 1282 m elevation, and two localities 8.1 km apart in Boualapha District, Khammouan Province, 980–1320 m, Laos.

Geographic Occurrence

Natural Resident: Laos

Endemic: Laos

Comment

In the Tylototriton asperrimus group according to the original publication. Yuan, Jiang, Lü, Yang, Nguyen, Nguyen, Jin, and Che, 2011, Zool. Res., Kunming, 32: 57–58, suggested that Tylototriton notialis is a junior synonym of Tylototrion asperrimus, but this conclusion was based on fewer data than the original recognition of the species. Raffaëlli, 2013, Urodeles du Monde, 2nd ed.: 186–187 provided a brief account, photograph, and map and documented the presence of this species in Vietnam, although only in provincial terms. Nishikawa, Matsui, and Nguyen, 2013, Curr. Herpetol., Kyoto, 32: 34–49, provided records from Nghe An Province, Vietnam (see below). Sparreboom, 2014, Salamanders Old World: 365–366, reviewed the biology, characteristics, distribution, reproduction, and conservation of the species. See brief account by Phimmachak, Aowphol, and Stuart, 2015, Zootaxa, 4006: 296–297. Bernardes, Le, Nguyen, Pham, Pham, Nguyen, Rödder, Bonkowski, and Ziegler, 2020, ZooKeys, 935: 121–164, provided a range map. Nishikawa, Sanamxay, Kurita, Hibino, Jono, and Nguyen, 2020, Acad. J. Biol., 42: 135–137, provided an upper elevation record. In the Tylototriton (Yaotriton) asperrimus species group of Poyarkov, Nguyen, and Arkhipov, 2021, Taprobanica, 10: 4–22, who discussed phylogenetics and reassigned the Vietnam record to a new species, Tylototriton thaiorum. Poyarkov, Nguyen, Popov, Geissler, Pawangkhanant, Neang, Suwannapoom, and Orlov, 2021, Russ. J. Herpetol., 28 (3A): 67, continued the inclusion of Vietnam within the range. In the Tylototriton asperrimus group of Lyu, Wang, Zeng, Zhou, Qi, Wan, Li, and Wang, 2021, Vert. Zool., Senckenberg, 71: 697–710, who discussed phylogenetics. Raffaëlli, 2022, Salamanders & Newts of the World: 288, provided an account, summarizing systematics, life history, population status, and distribution (including a polygon map). Stuart, Seateun, Sivongxay, and Phimmachak, 2025 "2024", in Wildlife Lao PDR: 20, briefly summarized literature, taxonomy, habitat, and range for Laos.

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For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist