Tylototriton panwaensis Grismer, Wood, Quah, Thura, Espinoza, and Murdoch, 2019

Class: Amphibia > Order: Caudata > Family: Salamandridae > Subfamily: Pleurodelinae > Genus: Tylototriton > Species: Tylototriton panwaensis

Tylototriton panwaensis Grismer, Wood, Quah, Thura, Espinoza, and Murdoch, 2019, J. Nat. Hist., London, 53: 483. Holotype: CAS 245418, by original designation. Type locality: "vicinity of Panwa Village, Panwa Township, Myitkyina District, Kachin State, Myanmar (25.60425° N, 98.37664°E, WGS84) at 2256 m in elevation". urn:lsid:zoobank.org:pub:BE851CE6-81BD-488C-A616-39A948A2B9C6 [Note: as of 15 July 2026 Zoobank continues to regard this name as unpublished; the date of publication therefore takes it date from the receipt of paper-printed copies — DRF]

Tylototriton joe Rao, Zeng, Zhu, and Ma in Rao, 2022 "2020", in Zhu and Rao (eds.), Atlas Wildlife SW China: 426. Holotype: KIZ-GLGS 16351; not available for examination according to Wang, Yu, Wu, Hou, Wang, Xiong, Ye, Duan, Li, LI, Jin, Yang, and Che, 2025, Zootaxa, 5725: 224–225, who suggested that a neotype designation may be required. Type locality: Pianma Town, Lushui County, Nujiang Lisu Autonomous Prefecture, Yunnan Province, China [from the Chinese]. Synonymy by  Che and Yu, 2026, Amph. Mt. Gaoligong: 6

Tylototriton (Tylototriton) joe — Raffaëlli, 2022, Salamanders & Newts of the World: 309.

Tylototriton (Tylototriton) panwaensis — Raffaëlli, 2022, Salamanders & Newts of the World: 316. 

Common Names

Panwa Crocodile Newt (Zug, 2022, Smithson. Contrib. Zool., 653: 7; Raffaëlli, 2022, Salamanders & Newts of the World: 309).

邦瓦疣螈 (Chinese: Che and Yu, 2026, Amph. Mt. Gaoligong: 6). 

Distribution

Northeastern Myanmar into western and northwestern Yunnan, China; a closely related population, possibly a distinct species is known from 304 km to the west at Lahe Township, Khandi District, Sagaing Region, and from the Kakhien Hillis of northern Myanmar. Also reported from Yingjiang and Tenchong, southwestern Yunnan, China. See comment.

Geographic Occurrence

Natural Resident: China, People's Republic of, Myanmar

Comment

 Zaw, Yin, Chit, and Hmwe, 2020, Univ. Mandalay Res. J., 11: 59–72, characterized the population from Sadung, Myitkyina District, Kachin State, Myanmar, via morphometrics and molecular markers. In the Tylototriton (Tylototriton) verrucosus species group of Poyarkov, Nguyen, and Arkhipov, 2021, Taprobanica, 10: 4–22, who discussed phylogenetics. In the Tylototriton verrucosus group of Lyu, Wang, Zeng, Zhou, Qi, Wan, Li, and Wang, 2021, Vert. Zool., Senckenberg, 71: 697–710, who discussed phylogenetics. Zug, 2022, Smithson. Contrib. Zool., 653: 7, discussed identification, habitat, and range in Myanmar. Liu, Hou, and Rao, 2022, Biodiversity Data J., 10 (e82707): 1–14, reported the species from Tenchong, Yunnan, China, and speculated that this species is a synonym of Tylototriton verrucosus, largely due to the latter's imprecise type locality; the authors suggested further work is needed inasmuch as the type locality of this species is very close to that of Tylototriton verrucosus, the type of the genus, and may be conspecific. Raffaëlli, 2022, Salamanders & Newts of the World: 309–310 (as Tylototriton joe) and 316 (as Tylototriton panwaensis), provided an account, summarizing systematics, life history, population status, and distribution (including a polygon map). Sánchez-Vialas, Copete-Mosquera, and Calvo-Revuelta, 2024, Zootaxa, 5457: 31, noted a specimen collected in the Kakhien Hillis of northern Myanmar and regarded specimens of Tylototriton verrucosus from this same region to be referrable to Tylototriton panwaensis. See brief account in Che and Yu, 2026, Amph. Mt. Gaoligong: 6, for Mount Gaoligong, Yunnan, China. 

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