Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Tylototriton panwaensis Grismer, Wood, Quah, Thura, Espinoza, and Murdoch, 2019

Class: Amphibia > Order: Caudata > Family: Salamandridae > Subfamily: Pleurodelinae > Genus: Tylototriton > Species: Tylototriton panwaensis

Tylototriton panwaensis Grismer, Wood, Quah, Thura, Espinoza, and Murdoch, 2019, J. Nat. Hist., London, 53: 483. Holotype: CAS 245418, by original designation. Type locality: "vicinity of Panwa Village, Panwa Township, Myitkyina District, Kachin State, Myanmar (25.60425° N, 98.37664°E, WGS84) at 2256 m in elevation". urn:lsid:zoobank.org:pub:BE851CE6-81BD-488C-A616-39A948A2B9C6 [Note: as of 12 Jan 2024 Zoobank regards this name as unpublished; the date of publication therefore takes it date from the receipt of paper-printed copies — DRF]

Tylototriton (Tylototriton) panwaensis — Raffaëlli, 2022, Salamanders & Newts of the World: 316.

Common Names

Panwa Crocodile Newt (Zug, 2022, Smithson. Contrib. Zool., 653: 7).

Distribution

Known from the vicinity of Panwa Town (Panwa Township) and Sadung (Myitkyina Township), Myitkyina District, Kachin State, Myanmar; a closely related population, possibly a distinct species is known from 304 km to the west at Lahe Township, Khandi District, Sagaing Region, and from the Kakhien Hillis of northern Myanmar. Also reported from Yingjiang and Tenchong, southwestern Yunnan, China. See comment.

Geographic Occurrence

Natural Resident: China, People's Republic of, Myanmar

Comment

Zaw, Yin, Chit, and Hmwe, 2020, Univ. Mandalay Res. J., 11: 59–72, characterized the population from Sadung, Myitkyina District, Kachin State, Myanmar, via morphometrics and molecular markers. In the Tylototriton (Tylototriton) verrucosus species group of Poyarkov, Nguyen, and Arkhipov, 2021, Taprobanica, 10: 4–22, who discussed phylogenetics. In the Tylototriton verrucosus group of Lyu, Wang, Zeng, Zhou, Qi, Wan, Li, and Wang, 2021, Vert. Zool., Senckenberg, 71: 697–710, who discussed phylogenetics. Zug, 2022, Smithson. Contrib. Zool., 653: 7, discussed identification, habitat, and range in Myanmar. Liu, Hou, and Rao, 2022, Biodiversity Data J., 10 (e82707): 1–14, reported the species from Tenchong, Yunnan, China, and speculated that this species is a synonym of Tylototriton verrucosus, largely due to the latter's imprecise type locality; the authors suggested further work is needed inasmuch as the type locality of this species is very close to that of Tylototriton verrucosus, the type of the genus, and may be conspecific. Raffaëlli, 2022, Salamanders & Newts of the World: 316, provided an account, summarizing systematics, life history, population status, and distribution (including a polygon map). Sánchez-Vialas, Copete-Mosquera, and Calvo-Revuelta, 2024, Zootaxa, 5457: 31, noted a specimen collected in the Kakhien Hillis of northern Myanmar and regarded specimens of Tylototriton verrucosus from this same region to be referrable to Tylototriton panwaensis.

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For additional information specific to China see Amphibia China