Tylototriton pulcherrimus Hou, Zhang, Li, and Lu, 2012

Class: Amphibia > Order: Caudata > Family: Salamandridae > Subfamily: Pleurodelinae > Genus: Tylototriton > Species: Tylototriton pulcherrimus

Tylototriton (Tylototriton) verrucosus pulcherrima Hou, Zhang, Li, and Lu in Hou, Li, and Lü, 2012, J. Huangshan Univ., 14: 63. Holotype: SYNU HM2012501-NT001, by original designation. Type locality: Luchun County, Yunnan, China. Synonymy by Nishikawa, Khonsue, Pomchote, and Matsui, 2013, Zootaxa, 3737: 277. Error in the gender of the subspecies name. Synonymy not accepted without comment by Bernardes, Le, Nguyen, Pham, Pham, Nguyen, Rödder, Bonkowski, and Ziegler, 2020, ZooKeys, 935: 121–164. 

Tylototriton pulcherrima —  Fei, Ye, and Jiang, 2012, Colored Atlas Chinese Amph. Distr.: 81. Incorrect gender of the species name. 

Tylototriton (Tylototriton) pulcherrima —  Raffaëlli, 2013, Urodeles du Monde, 2nd ed.: 183. Incorrect gender of the species name. 

Tylototriton pulcherrimus — Bernardes, Le, Nguyen, Pham, Pham, Nguyen, Rödder, Bonkowski, and Ziegler, 2020, ZooKeys, 935: 156. 

English Names

Hoanglien Mountain Crocodile Newt (Tylototriton verrucosus pulcherrimaHou, Li, and Lü, 2012, J. Huangshan Univ., 14: 63).

Huanglienshan Knobby Newt (Fei and Ye, 2016, Amph. China, 1: 272).

Hoang Lien Corcodile [sic] Newt (Poyarkov, Nguyen, Popov, Geissler, Pawangkhanant, Neang, Suwannapoom, and Orlov, 2021, Russ. J. Herpetol., 28 (3A): 67). 

Hoanglien Crocodile Newt (Raffaëlli, 2022, Salamanders & Newts of the World: 310).

   

Distribution

Southern Yunnan, China, and presumably extending along the Huanglian Mountains into adjacent northeastern Vietnam (Lai Chau Province); Long Phu and Phou Den Din in Laos but requiring confirmation. 

Comment

 Fei, Ye, and Jiang, 2012, Colored Atlas Chinese Amph. Distr.: 81, provided an account. Fei and Ye, 2016, Amph. China, 1: 272–277, provided account, photographs, and range map. Nishikawa, Khonsue, Pomchote, and Matsui, 2013, Zootaxa, 3737: 277, considered this species to be a synonym of Tylototriton shanjing on the basis of low genetic distance, but subsequent authors, mostly without comment, treated Tylototriton pulcherrimus as distinct. Hernandez and Hou, 2018, Bull. Soc. Herpetol. France, 165: 17–18, discussed the biogeography, conservation, ecology, and range in Yunnan and the likely range in Vietnam and Laos. Bernardes, Le, Nguyen, Pham, Pham, Nguyen, Rödder, Bonkowski, and Ziegler, 2020, ZooKeys, 935: 121–164, mapped the species as largely restricted to Yunnan, China, but extending to Myanmar borderlands; they treated Tylototriton pulcherrimus as distinct without comment but provided a table of comparisons that suggested that recognition was the conservative course at this point in time. In the Tylototriton (Tylototritonverrucosus species group of Poyarkov, Nguyen, and Arkhipov, 2021, Taprobanica, 10: 4–22, who discussed phylogenetics. In the Tylototriton verrucosus group of Lyu, Wang, Zeng, Zhou, Qi, Wan, Li, and Wang, 2021, Vert. Zool., Senckenberg, 71: 697–710, who discussed phylogenetics. Raffaëlli, 2022, Salamanders & Newts of the World: 310, provided an account, summarizing systematics, life history, population status, and distribution (including a polygon map). 

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