Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Tylototriton pulcherrimus Hou, Zhang, Li, and Lu, 2012

Class: Amphibia > Order: Caudata > Family: Salamandridae > Subfamily: Pleurodelinae > Genus: Tylototriton > Species: Tylototriton pulcherrimus

Tylototriton (Tylototriton) verrucosus pulcherrima Hou, Zhang, Li, and Lu in Hou, Li, and Lü, 2012, J. Huangshan Univ., 14: 63. Holotype: SYNU HM2012501-NT001, by original designation. Type locality: Luchun County, Yunnan, China. Synonymy by Nishikawa, Khonsue, Pomchote, and Matsui, 2013, Zootaxa, 3737: 277. Error in the gender of the subspecies name. Synonymy not accepted without comment by Bernardes, Le, Nguyen, Pham, Pham, Nguyen, Rödder, Bonkowski, and Ziegler, 2020, ZooKeys, 935: 121–164.

Tylototriton pulcherrima — Fei, Ye, and Jiang, 2012, Colored Atlas Chinese Amph. Distr.: 81. Incorrect gender of the species name.

Tylototriton (Tylototriton) pulcherrima — Raffaëlli, 2013, Urodeles du Monde, 2nd ed.: 183. Incorrect gender of the species name.

Tylototriton pulcherrimus — Bernardes, Le, Nguyen, Pham, Pham, Nguyen, Rödder, Bonkowski, and Ziegler, 2020, ZooKeys, 935: 156.

Common Names

Hoanglien Mountain Crocodile Newt (Tylototriton verrucosus pulcherrima: Hou, Li, and Lü, 2012, J. Huangshan Univ., 14: 63).

Huanglienshan Knobby Newt (Fei and Ye, 2016, Amph. China, 1: 272).

Hoang Lien Corcodile [sic] Newt (Poyarkov, Nguyen, Popov, Geissler, Pawangkhanant, Neang, Suwannapoom, and Orlov, 2021, Russ. J. Herpetol., 28 (3A): 67).

Hoanglien Crocodile Newt (Raffaëlli, 2022, Salamanders & Newts of the World: 310).

Distribution

Southern Yunnan, China, and presumably extending along the Huanglian Mountains into adjacent northeastern Vietnam (Lai Chau Province); Long Phu and Phou Den Din in Laos but requiring confirmation.

Geographic Occurrence

Natural Resident: China, People's Republic of

Likely/Controversially Present: Laos, Vietnam

Endemic: China, People's Republic of

Comment

Fei, Ye, and Jiang, 2012, Colored Atlas Chinese Amph. Distr.: 81, provided an account. Fei and Ye, 2016, Amph. China, 1: 272–277, provided account, photographs, and range map. Nishikawa, Khonsue, Pomchote, and Matsui, 2013, Zootaxa, 3737: 277, considered this species to be a synonym of Tylototriton shanjing on the basis of low genetic distance, but subsequent authors, mostly without comment, treated Tylototriton pulcherrimus as distinct. Hernandez and Hou, 2018, Bull. Soc. Herpetol. France, 165: 17–18, discussed the biogeography, conservation, ecology, and range in Yunnan and the likely range in Vietnam and Laos. Bernardes, Le, Nguyen, Pham, Pham, Nguyen, Rödder, Bonkowski, and Ziegler, 2020, ZooKeys, 935: 121–164, mapped the species as largely restricted to Yunnan, China, but extending to Myanmar borderlands; they treated Tylototriton pulcherrimus as distinct without comment but provided a table of comparisons that suggested that recognition was the conservative course at this point in time. In the Tylototriton (Tylototriton) verrucosus species group of Poyarkov, Nguyen, and Arkhipov, 2021, Taprobanica, 10: 4–22, who discussed phylogenetics. In the Tylototriton verrucosus group of Lyu, Wang, Zeng, Zhou, Qi, Wan, Li, and Wang, 2021, Vert. Zool., Senckenberg, 71: 697–710, who discussed phylogenetics. Raffaëlli, 2022, Salamanders & Newts of the World: 310, provided an account, summarizing systematics, life history, population status, and distribution (including a polygon map). Not included in the fauna of Laos by Stuart, Seateun, Sivongxay, and Phimmachak, 2025 "2024", in Wildlife Lao PDR: 17–47.

External links:

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For access to general information see Wikipedia
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To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist
For additional information specific to China see Amphibia China