Tylototriton shanjing Nussbaum, Brodie, and Yang, 1995

Class: Amphibia > Order: Caudata > Family: Salamandridae > Subfamily: Pleurodelinae > Genus: Tylototriton > Species: Tylototriton shanjing

Tylototriton shanjing Nussbaum, Brodie, and Yang, 1995, Herpetologica, 51: 265. Holotype: KIZ II 0731 V.27, by original designation. Type locality: "Dingpa, 2150 m elevation, Jingdong County, Yunnan Province, People's Republic of China". 

Tylototriton verrucosus shanjing — Yang, 2008, in Yang and Rao (ed.), Amph. Rept. Yunnan: 19.

Tylototriton (Tylototriton) shanjing — Dubois and Raffaëlli, 2009, Alytes, 26: 68.

English Names

Red Knobby Newt (Fei and Ye, 2016, Amph. China, 1: 275). 

Yunnan Newt (Sparreboom, 2014, Salamanders Old World: 369). 

Mandarin Salamander (Sparreboom, 2014, Salamanders Old World: 369). 


Western, central, and southern Yunnan, China, 1000 to 2000 m elevation; probably to be found in northern Thailand. See comment.  


Tylototriton shanjing had been placed in the synonymy of Tylototriton verrucosus by Zhang, Rao, Yu, and Yang, 2007, Zool. Res., Kunming, 28: 430–436, on the basis of minimal sequence divergence; this synonymy disputed implicitly by Dubois and Raffaëlli, 2009, Alytes, 26: 68, and explicitly by Stuart, Phimmachak, Sivongxay, and Robichaud, 2010, Zootaxa, 2650: 28. See brief account by Zhao and Yang, 1997, Amph. Rept. Hengduan Mountains Region: 34-35. In the Tylototriton verrucosus group of Fei, Ye, Huang, Jiang, and Xie, 2005, in Fei et al. (eds.), Illust. Key Chinese Amph.: 42 (although they only addressed Chinese species). Fei, Hu, Ye, and Huang, 2006, Fauna Sinica, Amph. 1: 272-276, provided an account and range map. Raffaëlli, 2013, Urodeles du Monde, 2nd ed.: 181-182, provided a brief account, figure, and map. Yang, 2008, in Yang and Rao (ed.), Amph. Rept. Yunnan: 18-19, provided a brief account for Yunnan, China. See statement of geographic range, habitat, and conservation status in Stuart, Hoffmann, Chanson, Cox, Berridge, Ramani, and Young, 2008, Threatened Amph. World: 642. Fei, Ye, and Jiang, 2010, Colored Atlas of Chinese Amph.: 77. Fei, Ye, and Jiang, 2012, Colored Atlas Chinese Amph. Distr.: 86, provided an account.  Zhao, Rao, Liu, Li, and Yuan, 2012, J. W. China Forest. Sci., 41: 85–89, provided molecular evidence for the distinctiveness of this species and it sister-taxon relationship with Tylototriton daweishanensis.  Raffaëlli, 2013, Urodeles du Monde, 2nd ed.: 183, provided a brief account, photograph, and map. Records for Laos now have been referred to Tylototriton podichthysSparreboom, 2014, Salamanders Old World: 369–371, reviewed the biology, characteristics, distribution, reproduction, and conservation of the species. Khatiwada, Wang, Ghimire, Vasudevan, Paudel, and Jiang, 2015, Asian Herpetol. Res., 6: 251, provided molecular evidence that suggests that Tylototriton shanjing is composed of two lineages, one of which is nested within Tylototriton verrucosusFei and Ye, 2016, Amph. China, 1: 272–277, provided accounts (as Tylototriton shanjing and, without comment Tylototriton pulcherrima [sic], photographs, and range map, although given the results of  Nishikawa, Khonsue, Pomchote, and Matsui, 2013, Zootaxa, 3737: 277, one might expect nominal Tylototriton shanjing to be found to be a species complex. See comment under Tylototriton pulcherrimus. In the Tylototriton (Tylototritonverrucosus species group of Poyarkov, Nguyen, and Arkhipov, 2021, Taprobanica, 10: 4–22, who discussed phylogenetics. In the Tylototriton verrucosus group of Lyu, Wang, Zeng, Zhou, Qi, Wan, Li, and Wang, 2021, Vert. Zool., Senckenberg, 71: 697–710, who discussed phylogenetics. Raffaëlli, 2022, Salamanders & Newts of the World: 306–308, provided an account, summarizing systematics, life history, population status, and distribution (including a polygon map). 

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