Tylototriton shanjing Nussbaum, Brodie, and Yang, 1995

Class: Amphibia > Order: Caudata > Family: Salamandridae > Subfamily: Pleurodelinae > Genus: Tylototriton > Species: Tylototriton shanjing

Tylototriton shanjing Nussbaum, Brodie, and Yang, 1995, Herpetologica, 51: 265. Holotype: KIZ II 0731 V.27, by original designation. Type locality: "Dingpa, 2150 m elevation, Jingdong County, Yunnan Province, People's Republic of China". 

Tylototriton verrucosus shanjing — Yang, 2008, in Yang and Rao (ed.), Amph. Rept. Yunnan: 19.

Tylototriton (Tylototriton) shanjing — Dubois and Raffaëlli, 2009, Alytes, 26: 68.

English Names

Red Knobby Newt (Fei and Ye, 2016, Amph. China, 1: 275). 

Yunnan Newt (Sparreboom, 2014, Salamanders Old World: 369). 

Mandarin Salamander (Sparreboom, 2014, Salamanders Old World: 369). 

Distribution

Western and central Yunnan, China, extending to northeastern and eastern Myanmar borderlands, 1000 to 2000 m elevation. See comment.  

Comment

Tylototriton shanjing had been placed in the synonymy of Tylototriton verrucosus by Zhang, Rao, Yu, and Yang, 2007, Zool. Res., Kunming, 28: 430–436, on the basis of minimal sequence divergence; this synonymy disputed implicitly by Dubois and Raffaëlli, 2009, Alytes, 26: 68, and explicitly by Stuart, Phimmachak, Sivongxay, and Robichaud, 2010, Zootaxa, 2650: 28. See brief account by Zhao and Yang, 1997, Amph. Rept. Hengduan Mountains Region: 34-35. In the Tylototriton verrucosus group of Fei, Ye, Huang, Jiang, and Xie, 2005, in Fei et al. (eds.), Illust. Key Chinese Amph.: 42 (although they only addressed Chinese species). Fei, Hu, Ye, and Huang, 2006, Fauna Sinica, Amph. 1: 272-276, provided an account and range map. Raffaëlli, 2013, Urodeles du Monde, 2nd ed.: 181-182, provided a brief account, figure, and map. Yang, 2008, in Yang and Rao (ed.), Amph. Rept. Yunnan: 18-19, provided a brief account for Yunnan, China. See statement of geographic range, habitat, and conservation status in Stuart, Hoffmann, Chanson, Cox, Berridge, Ramani, and Young, 2008, Threatened Amph. World: 642. Fei, Ye, and Jiang, 2010, Colored Atlas of Chinese Amph.: 77. Fei, Ye, and Jiang, 2012, Colored Atlas Chinese Amph. Distr.: 86, provided an account.  Zhao, Rao, Liu, Li, and Yuan, 2012, J. W. China Forest. Sci., 41: 85–89, provided molecular evidence for the distinctiveness of this species and it sister-taxon relationship with Tylototriton daweishanensis.  Raffaëlli, 2013, Urodeles du Monde, 2nd ed.: 183, provided a brief account, photograph, and map. Records for Laos now have been referred to Tylototriton podichthysSparreboom, 2014, Salamanders Old World: 369–371, reviewed the biology, characteristics, distribution, reproduction, and conservation of the species. Khatiwada, Wang, Ghimire, Vasudevan, Paudel, and Jiang, 2015, Asian Herpetol. Res., 6: 251, provided molecular evidence that suggests that Tylototriton shanjing is composed of two lineages, one of which is nested within Tylototriton verrucosusFei and Ye, 2016, Amph. China, 1: 272–277, provided accounts (as Tylototriton shanjing and, without comment Tylototriton pulcherrima [sic], photographs, and range map, although given the results of  Nishikawa, Khonsue, Pomchote, and Matsui, 2013, Zootaxa, 3737: 277, one might expect nominal Tylototriton shanjing to be found to be a species complex. See comment under Tylototriton pulcherrimus. In the Tylototriton (Tylototritonverrucosus species group of Poyarkov, Nguyen, and Arkhipov, 2021, Taprobanica, 10: 4–22, who discussed phylogenetics. 

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