Tylototriton shanorum Nishikawa, Matsui, and Rao, 2014

Class: Amphibia > Order: Caudata > Family: Salamandridae > Subfamily: Pleurodelinae > Genus: Tylototriton > Species: Tylototriton shanorum

Tylototriton (Tylototriton) shanorum Nishikawa, Matsui, and Rao, 2014, Nat. Hist. Bull. Siam Soc., 60: 16. Holotype: CAS 230940, by original designation. Type locality: "Taunggyi Township, Shan State, Myanmar (20° 48′ 28.5″ N, 97°02′ 45.1″ E, 1457 m asl)". 

Tylototriton ngarsuensis Grismer, Wood, Quah, Thura, Espinoza, Grismer, Murdoch, and Lin, 2018, Zootaxa, 4500: 562. Holotype: LSUHC 13762, by original designation. Type locality: "Baw Hto Chang in Ngar Su Village, Ywangan Township, Taunggyi District, Shan State, Myanmar (21.15364° N, 96.43660°E WGS84) at 1212 m in elevation." Zoobank publication registration: 3518D551-CA4B-43A4-B6D3-8A159E2398E1. Synonymy by Onishi, Kyaw, Nishikawa, Myint, and Thwe, 2025, Zootaxa, 5620: 401. 

Common Names

Shan Crocodile Newt (Zug, 2022, Smithson. Contrib. Zool., 653: 7). 

Taunggyi Crocodile Newt (Raffaëlli, 2022, Salamanders & Newts of the World: 316).

Ywangan Crocodile Newt (Tylototriton ngarsuensis [no longer recognized]: Grismer, Wood, Quah, Thura, Espinoza, Grismer, Murdoch, and Lin, 2018, Zootaxa, 4500: 562). 

Ayeyarwady Crocodile Newt (Tylototriton ngarsuensis [no longer recognized]: Zug, 2022, Smithson. Contrib. Zool., 653: 7) 

Distribution

Taunggyi, Kalaw, Pindaya, Nyaungshwe, Pinlaung, and Ywnagan townships, Shan state, central Myanmar. 

Geographic Occurrence

Natural Resident: Myanmar

Endemic: Myanmar

Comment

Previously confused with Tylototriton verrucosus, but actually distant from that species according to an ML tree of ND2 in the original publication, where an unnamed Nepal population was also discussed. See comment under Tylototriton himalayanus.  Grismer, Wood, Quah, Thura, Espinoza, Grismer, Murdoch, and Lin, 2018, Zootaxa, 4500: 553–573, discussed in detail the morphology and phylogenetic relations of this species. Bernardes, Le, Nguyen, Pham, Pham, Nguyen, Rödder, Bonkowski, and Ziegler, 2020, ZooKeys, 935: 121–164, mapped nominal Tylototriton shanorum and Tylototriton ngarsuensis. Onishi, Kyaw, Aung, Myint, Thwe, and Nishikawa, 2020, Nat. Hist. Bull. Siam Soc., 64: 47–50, provided new localities and discussed the range. In the Tylototriton (Tylototritonverrucosus species group of Poyarkov, Nguyen, and Arkhipov, 2021, Taprobanica, 10: 4–22, who discussed phylogenetics. In the Tylototriton verrucosus group of Lyu, Wang, Zeng, Zhou, Qi, Wan, Li, and Wang, 2021, Vert. Zool., Senckenberg, 71: 697–710, who discussed phylogenetics. Zug, 2022, Smithson. Contrib. Zool., 653: 7, discussed identification, habitat, and range of nominal Tylototriton ngarsuensis and Tylototritonin shanorum in Myanmar. Raffaëlli, 2022, Salamanders & Newts of the World: 316–318, provided  accounts of Tylototriton shanorum and Tylototriton ngarsuensis, summarizing systematics, life history, population status, and distribution (including a polygon map). Onishi, Kyaw, Nishikawa, Myint, and Thwe, 2025, Zootaxa, 5620: 401–425, discussed landscape genetics and taxonomy.  

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