Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Tylototriton shanorum Nishikawa, Matsui, and Rao, 2014

Class: Amphibia > Order: Caudata > Family: Salamandridae > Subfamily: Pleurodelinae > Genus: Tylototriton > Species: Tylototriton shanorum

Tylototriton (Tylototriton) shanorum Nishikawa, Matsui, and Rao, 2014, Nat. Hist. Bull. Siam Soc., 60: 16. Holotype: CAS 230940, by original designation. Type locality: "Taunggyi Township, Shan State, Myanmar (20° 48′ 28.5″ N, 97°02′ 45.1″ E, 1457 m asl)".

Tylototriton ngarsuensis Grismer, Wood, Quah, Thura, Espinoza, Grismer, Murdoch, and Lin, 2018, Zootaxa, 4500: 562. Holotype: LSUHC 13762, by original designation. Type locality: "Baw Hto Chang in Ngar Su Village, Ywangan Township, Taunggyi District, Shan State, Myanmar (21.15364° N, 96.43660°E WGS84) at 1212 m in elevation." Zoobank publication registration: 3518D551-CA4B-43A4-B6D3-8A159E2398E1. Synonymy by Onishi, Kyaw, Nishikawa, Myint, and Thwe, 2025, Zootaxa, 5620: 401.

Common Names

Shan Crocodile Newt (Zug, 2022, Smithson. Contrib. Zool., 653: 7).

Taunggyi Crocodile Newt (Raffaëlli, 2022, Salamanders & Newts of the World: 316).

Ywangan Crocodile Newt (Tylototriton ngarsuensis [no longer recognized]: Grismer, Wood, Quah, Thura, Espinoza, Grismer, Murdoch, and Lin, 2018, Zootaxa, 4500: 562).

Ayeyarwady Crocodile Newt (Tylototriton ngarsuensis [no longer recognized]: Zug, 2022, Smithson. Contrib. Zool., 653: 7)

Distribution

Taunggyi, Kalaw, Pindaya, Nyaungshwe, Pinlaung, and Ywnagan townships, Shan state, central Myanmar.

Geographic Occurrence

Natural Resident: Myanmar

Endemic: Myanmar

Comment

Previously confused with Tylototriton verrucosus, but actually distant from that species according to an ML tree of ND2 in the original publication, where an unnamed Nepal population was also discussed. See comment under Tylototriton himalayanus. Grismer, Wood, Quah, Thura, Espinoza, Grismer, Murdoch, and Lin, 2018, Zootaxa, 4500: 553–573, discussed in detail the morphology and phylogenetic relations of this species. Bernardes, Le, Nguyen, Pham, Pham, Nguyen, Rödder, Bonkowski, and Ziegler, 2020, ZooKeys, 935: 121–164, mapped nominal Tylototriton shanorum and Tylototriton ngarsuensis. Onishi, Kyaw, Aung, Myint, Thwe, and Nishikawa, 2020, Nat. Hist. Bull. Siam Soc., 64: 47–50, provided new localities and discussed the range. In the Tylototriton (Tylototriton) verrucosus species group of Poyarkov, Nguyen, and Arkhipov, 2021, Taprobanica, 10: 4–22, who discussed phylogenetics. In the Tylototriton verrucosus group of Lyu, Wang, Zeng, Zhou, Qi, Wan, Li, and Wang, 2021, Vert. Zool., Senckenberg, 71: 697–710, who discussed phylogenetics. Zug, 2022, Smithson. Contrib. Zool., 653: 7, discussed identification, habitat, and range of nominal Tylototriton ngarsuensis and Tylototritonin shanorum in Myanmar. Raffaëlli, 2022, Salamanders & Newts of the World: 316–318, provided accounts of Tylototriton shanorum and Tylototriton ngarsuensis, summarizing systematics, life history, population status, and distribution (including a polygon map). Onishi, Kyaw, Nishikawa, Myint, and Thwe, 2025, Zootaxa, 5620: 401–425, discussed landscape genetics and taxonomy.

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For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist