Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Tylototriton uyenoi Nishikawa, Khonsue, Pomchote, and Matsui, 2013

Class: Amphibia > Order: Caudata > Family: Salamandridae > Subfamily: Pleurodelinae > Genus: Tylototriton > Species: Tylototriton uyenoi

Tylototriton (Tylototriton) uyenoi Nishikawa, Khonsue, Pomchote, and Matsui, 2013, Zootaxa, 3737: 272. Holotype: KUHE 19147, by original designation. Type locality: "Phuping Rajanives Palace, Doi Suthep, Chiang Mai Province, Thailand (18 48’16’’ N, 98 54’9’’ E, 1436 m asl)".

Common Names

Uéno's Knobby Newt (Sparreboom, 2014, Salamanders Old World: 372).

Chiang Mai Corcodile [sic] Newt (Poyarkov, Nguyen, Popov, Geissler, Pawangkhanant, Neang, Suwannapoom, and Orlov, 2021, Russ. J. Herpetol., 28 (3A): 67).

Chiang Mai Crocodile Newt (Raffaëlli, 2022, Salamanders & Newts of the World: 313).

Distribution

Chiang Mai Province (Doi Ang Khang, Doi Chang Kien, Doi Inthanon, Doi Pui, Chiang Mai, and Doi Suthep), northwestern Thailand and Khao Laem National Park, Kanchanaburi Province, western Thailand at 1507 m elevation; likely extending into adjacent Myanmar.

Geographic Occurrence

Natural Resident: Thailand

Likely/Controversially Present: Myanmar

Endemic: Thailand

Comment

Previously confused with Tylototriton verrucosus according to the original publication. Raffaëlli, 2013, Urodeles du Monde, 2nd ed.: 182-183, provided a brief account, photograph, and map. Sparreboom, 2014, Salamanders Old World: 372–373, reviewed the biology, characteristics, distribution, reproduction, and conservation of the species. Niyomwan, Srisom, and Pawangkhanant, 2019, Field Guide Amph. Thailand: 96–97, provided a brief account (photograph, habitat, and range) for Thailand (in Thai). Hernandez, Escoriza, Pomchote, and Hou, 2019, Herpetol. Bull., London, 147: 15–18, provided a record for Tak Province, northwestern Thailand, and discussed the range and habitat. Bernardes, Le, Nguyen, Pham, Pham, Nguyen, Rödder, Bonkowski, and Ziegler, 2020, ZooKeys, 935: 121–164 , provided a range map that suggested the presence of this species in Myanmar and Vietnam, although the provenance of specimens was not provided. Hernandez and Pomchote, 2020, Bull. Soc. Herpetol. France, 175: 64–67, provided a record from Khao Laem National Park, Kanchanaburi province, western Thailand. Makchai, Chuaynkern, Safoowong, Chuachat, and Cota, 2020, Amph. N. Thailand: 188–189, provided photographs, a brief account for Thailand, and a range map. In the Tylototriton (Tylototriton) verrucosus species group of Poyarkov, Nguyen, and Arkhipov, 2021, Taprobanica, 10: 4–22, who discussed phylogenetics. Rongchapho, Nampochai, Makchai, and Cota, Thailand Nat. Hist. Mus. J., 15: 59–64, reported on morphology and distribution within Chiang Mai Province, Thailand. In the Tylototriton verrucosus group of Lyu, Wang, Zeng, Zhou, Qi, Wan, Li, and Wang, 2021, Vert. Zool., Senckenberg, 71: 697–710, who discussed phylogenetics. Population from Tak Province, Thailand, is now assigned to Tylototriton umphangensis. Raffaëlli, 2022, Salamanders & Newts of the World: 313–314, provided an account, summarizing systematics, life history, population status, and distribution (including a polygon map).

External links:

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For access to general information see Wikipedia
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For images search CalPhoto Images and Google Images
To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist