Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Tylototriton vietnamensis Böhme, Schöttler, Nguyen, and Köhler, 2005

Class: Amphibia > Order: Caudata > Family: Salamandridae > Subfamily: Pleurodelinae > Genus: Tylototriton > Species: Tylototriton vietnamensis

Tylototriton vietnamensis Böhme, Schöttler, Nguyen, and Köhler, 2005, Salamandra, 41: 215. Holotype: ZFMK 80637, by original designation. Type locality: "vicinity of Dong Vanh Village, Luc Son Commune, Luc Nam District, Bac Giang Province, northern Vietnam (21° 12´ N, 106° 40´ E, approximately 250-300 m a.s.l.)".

Tylototriton (Yaotriton) vietnamensis — Dubois and Raffaëlli, 2009, Alytes, 26: 68.

Yaotriton vietnamensis — Fei, Ye, and Jiang, 2012, Colored Atlas Chinese Amph. Distr.: 87, by implication; Dubois, Ohler, and Pyron, 2021, Megataxa, 5: 408.

Common Names

Vietnamese Knobby Newt (Sparreboom, 2014, Salamanders Old World: 378).

Vietnamese Crocodile Newt (Sparreboom, 2014, Salamanders Old World: 378).

Bac Giang Crocodile Newt (Raffaëlli, 2022, Salamanders & Newts of the World: 297).

Distribution

Northern and north-central Vietnam in Phu Tho, Cao Bang, Bac Giang, and Quang Ninh provinces; Khammouan Province, Laos.

Geographic Occurrence

Natural Resident: Vietnam

Endemic: Vietnam

Comment

See comments under Tylototriton and Tylototriton asperrimus. Raffaëlli, 2007, Les Urodèles du Monde: 148–149, provided a brief account, figure, and map. Nguyen, Nguyen, Hồ, Le, and Nguyen, 2009, Tạp chí Công nghệ Sinh học, 7: 325–333, discussed the species and its distinctiveness from Tylototriton hainanensis. Stuart, Phimmachak, Sivongxay, and Robichaud, 2010, Zootaxa, 2650: 30, suggested that nominal Tylototriton vietnamensis may include more than one species (Tylototriton zieglieri being one of the recent segregates—DRF). Bernardes, Rödder, Nguyen, Pham, Nguyen, and Ziegler, 2013, J. Nat. Hist., London, 47: 1161–1175, discussed habitat and potential distribution. Hecht, Pham, Nguyen, Nguyen, Bonkowski, and Ziegler, 2013, Biodiversity J., 4: 507–552, reported a record from Tay Yen Tu Nature Reserve, Bac Giang Province, northeastern Vietnam and discussed the range. Sparreboom, 2014, Salamanders Old World: 378–379, reviewed the biology, characteristics, distribution, reproduction, and conservation of the species. The sister taxon of Tylototriton panhai according to Phimmachak, Aowphol, and Stuart, 2015, Zootaxa, 4006: 293, who provided a mtDNA tree of the species of Tylototriton. Bernardes, Pham, Nguyen, Le, Bonkowski, and Ziegler, 2017, Salamandra, 53: 451–457, reported on morphometrics and ecology of a population in Dong Son–Ky Thuong Nature Reserve, Dong Son Commune, Hoanh Bo District, Quang Ninh Province, Vietnam. In the Tylototriton (Yaotriton) vietnamensis species group of Poyarkov, Nguyen, and Arkhipov, 2021, Taprobanica, 10: 4–22, who discussed phylogenetics. In the Tylototriton asperrimus group of Lyu, Wang, Zeng, Zhou, Qi, Wan, Li, and Wang, 2021, Vert. Zool., Senckenberg, 71: 697–710, who discussed phylogenetics. Raffaëlli, 2022, Salamanders & Newts of the World: 297–298, provided an account, summarizing systematics, life history, population status, and distribution (including a polygon map).

External links:

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To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist