Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Tylototriton wenxianensis Fei, Ye, and Yang, 1984

Class: Amphibia > Order: Caudata > Family: Salamandridae > Subfamily: Pleurodelinae > Genus: Tylototriton > Species: Tylototriton wenxianensis

Tylototriton asperrimus wenxianensis Fei, Ye, and Yang, 1984, Acta Zool. Sinica, 30: 89. Holotype: CIB 638164, by original designation. Type locality: "Wenxian [= Wen County], Gansu [Province], alt. 946 m", China.

Tylototriton asperrimus pingwuensis Deng and Yu, 1984, Acta Herpetol. Sinica, Chengdu, N.S.,, 3 (2): 75. Holotype: KIZ 74005, by original designation. Type locality: "Duiwoliang, Pingwu, Sichuan, alt. 1400 m", China. Synonymy by Ye, Fei, and Hu, 1993, Rare and Economic Amph. China: 80, following Zhao, Hu, Jiang, and Yang, 1988, Studies on Chinese Salamanders: 63, who considered it likely synonymous.

Pleurodeles (Tylototrion) asperrimus wenxianensis — Risch, 1985, J. Bengal Nat. Hist. Soc., N.S.,, 4: 142.

Echinotriton asperrimus wenxianensis — Zhao and Adler, 1993, Herpetol. China: 112. 1

Tylototriton wenxianensis — Ye, Fei, and Hu, 1993, Rare and Economic Amph. China: 80.

Tylototriton (Yaotriton) wenxianensis — Dubois and Raffaëlli, 2009, Alytes, 26: 68.

Tylototriton wenxianensis wenxianensis — Chen, Wang, and Tao, 2010, Acta Zootaxon. Sinica, 35: 666; by implication.

Yaotriton wenxianensis — Fei, Ye, and Jiang, 2012, Colored Atlas Chinese Amph. Distr.: 94; Dubois, Ohler, and Pyron, 2021, Megataxa, 5: 408.

Common Names

Wenxian Knobby Salamander (Fei, 1999, Atlas Amph. China: 42).

Wenxian Knobby Newt (Fei and Ye, 2001, Color Handbook Amph. Sichuan: 96).

Wenxian Crocodile Newt (Raffaëlli, 2022, Salamanders & Newts of the World: 289).

Distribution

Chongqing (Fengjie, Wenzhou, and Yunyang), Gansu (Wenxian), Guizhou (Dafang, Leishan, Suiyang, and Zunyi), and Sichuan (Jiange, Leishan, Pingwu, Qingchuan, and Wangcang), China, 940 to 1400 m elevation. See comment.

Geographic Occurrence

Natural Resident: China, People's Republic of

Endemic: China, People's Republic of

Comment

See comments under Tylototriton. See accounts by Ye, Fei, and Hu, 1993, Rare and Economic Amph. China 81, and Fei, 1999, Atlas Amph. China: 42. Fei and Ye, 2001, Color Handbook Amph. Sichuan: 96, provided a brief account and illustration. In the Tylototriton asperrimus group of Fei, Ye, Huang, Jiang, and Xie, 2005, in Fei et al. (eds.), Illust. Key Chinese Amph.: 42 (although they only addressed Chinese species). Fei, Hu, Ye, and Huang, 2006, Fauna Sinica, Amph. 1: 265-268. Raffaëlli, 2013, Urodeles du Monde, 2nd ed.: 187, provided a brief account, photograph, and map. See photograph, map, description of geographic range and habitat, and conservation status in Stuart, Hoffmann, Chanson, Cox, Berridge, Ramani, and Young, 2008, Threatened Amph. World: 607. Fei, Ye, and Jiang, 2010, Colored Atlas of Chinese Amph.: 74-75, provided a brief account including photographs of specimens and habitat. Fei, Ye, and Jiang, 2012, Colored Atlas Chinese Amph. Distr.: 94–95, provided an account, photographs, and a map. Sparreboom, 2014, Salamanders Old World: 380–381, reviewed the biology, characteristics, distribution, reproduction, and conservation of the species. Fei and Ye, 2016, Amph. China, 1: 306–308, provided an account (as Yaotriton wenxianensis), photographs, and range map. See Yao and Gong, 2012, Amph. Rept. Gansu: 26–27, who provided a brief account and photograph, as Echinotriton asperrimus. Bernardes, Le, Nguyen, Pham, Pham, Nguyen, Rödder, Bonkowski, and Ziegler, 2020, ZooKeys, 935: 121–164, provided a range map, treating the populations from northern Guangxi, central Guizhou, Chonquing and Hubei, China as Tylototriton cf. wenxianensis, implying a species complex. In the Tylototriton (Yaotriton) wenxianensis species group of Poyarkov, Nguyen, and Arkhipov, 2021, Taprobanica, 10: 4–22, who discussed phylogenetics. In the Tylototriton asperrimus group of Lyu, Wang, Zeng, Zhou, Qi, Wan, Li, and Wang, 2021, Vert. Zool., Senckenberg, 71: 697–710, who discussed phylogenetics. Raffaëlli, 2022, Salamanders & Newts of the World: 289–291, provided an account, summarizing systematics, life history, population status, and distribution (including a polygon map).

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For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist
For additional information specific to China see Amphibia China