Tylototriton wenxianensis Fei, Ye, and Yang, 1984

Class: Amphibia > Order: Caudata > Family: Salamandridae > Subfamily: Pleurodelinae > Genus: Tylototriton > Species: Tylototriton wenxianensis

Tylototriton asperrimus wenxianensis Fei, Ye, and Yang, 1984, Acta Zool. Sinica, 30: 89. Holotype: CIB 638164, by original designation. Type locality: "Wenxian [= Wen County], Gansu [Province], alt. 946 m", China.

Tylototriton asperrimus pingwuensis Deng and Yu, 1984, Acta Herpetol. Sinica, Chengdu, N.S.,, 3 (2): 75. Holotype: KIZ 74005, by original designation. Type locality: "Duiwoliang, Pingwu, Sichuan, alt. 1400 m", China. Synonymy by Ye, Fei, and Hu, 1993, Rare and Economic Amph. China: 80, following Zhao, Hu, Jiang, and Yang, 1988, Studies on Chinese Salamanders: 63, who considered it likely synonymous.

Pleurodeles (Tylototrion) asperrimus wenxianensisRisch, 1985, J. Bengal Nat. Hist. Soc., N.S.,, 4: 142.

Echinotriton asperrimus wenxianensisZhao and Adler, 1993, Herpetol. China: 112. 1

Tylototriton wenxianensisYe, Fei, and Hu, 1993, Rare and Economic Amph. China: 80.

Tylototriton (Yaotriton) wenxianensisDubois and Raffaëlli, 2009, Alytes, 26: 68.

Tylototriton wenxianensis wenxianensisChen, Wang, and Tao, 2010, Acta Zootaxon. Sinica, 35: 666; by implication.

Yaotriton wenxianensis — Fei, Ye, and Jiang, 2012, Colored Atlas Chinese Amph. Distr.: 94; Dubois, Ohler, and Pyron, 2021, Megataxa, 5: 408.

English Names

Wenxian Knobby Salamander (Fei, 1999, Atlas Amph. China: 42).

Wenxian Knobby Newt (Fei and Ye, 2001, Color Handbook Amph. Sichuan: 96).


Chongqing (Fengjie, Wenzhou, and Yunyang), Gansu (Wenxian), Guizhou (Dafang, Leishan, Suiyang, and Zunyi), and Sichuan (Jiange, Leishan, Pingwu, Qingchuan, and Wangcang), China, 940 to 1400 m elevation. See comment. 


See comments under Tylototriton. See accounts by Ye, Fei, and Hu, 1993, Rare and Economic Amph. China 81, and Fei, 1999, Atlas Amph. China: 42. Fei and Ye, 2001, Color Handbook Amph. Sichuan: 96, provided a brief account and illustration. In the Tylototriton asperrimus group of Fei, Ye, Huang, Jiang, and Xie, 2005, in Fei et al. (eds.), Illust. Key Chinese Amph.: 42 (although they only addressed Chinese species). Fei, Hu, Ye, and Huang, 2006, Fauna Sinica, Amph. 1: 265-268. Raffaëlli, 2013, Urodeles du Monde, 2nd ed.: 187, provided a brief account, photograph, and map. See photograph, map, description of geographic range and habitat, and conservation status in Stuart, Hoffmann, Chanson, Cox, Berridge, Ramani, and Young, 2008, Threatened Amph. World: 607. Fei, Ye, and Jiang, 2010, Colored Atlas of Chinese Amph.: 74-75, provided a brief account including photographs of specimens and habitat. Fei, Ye, and Jiang, 2012, Colored Atlas Chinese Amph. Distr.: 94–95, provided an account, photographs, and a map. Sparreboom, 2014, Salamanders Old World: 380–381, reviewed the biology, characteristics, distribution, reproduction, and conservation of the species. Fei and Ye, 2016, Amph. China, 1: 306–308, provided an account (as Yaotriton wenxianensis), photographs, and range map. See Yao and Gong, 2012, Amph. Rept. Gansu: 26–27, who provided a brief account and photograph, as Echinotriton asperrimusBernardes, Le, Nguyen, Pham, Pham, Nguyen, Rödder, Bonkowski, and Ziegler, 2020, ZooKeys, 935: 121–164, provided a range map, treating the populations from northern Guangxi, central Guizhou, Chonquing and Hubei, China as Tylototriton cf. wenxianensis, implying a species complex. In the Tylototriton (Yaotritonwenxianensis species group of Poyarkov, Nguyen, and Arkhipov, 2021, Taprobanica, 10: 4–22, who discussed phylogenetics. In the Tylototriton asperrimus group of Lyu, Wang, Zeng, Zhou, Qi, Wan, Li, and Wang, 2021, Vert. Zool., Senckenberg, 71: 697–710, who discussed phylogenetics. 

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