Bufotes pseudoraddei (Mertens, 1971)

Class: Amphibia > Order: Anura > Family: Bufonidae > Genus: Bufotes > Species: Bufotes pseudoraddei

Bufo viridis pseudoraddei Mertens, 1971, Senckenb. Biol., 52: 8, and fig. 3. Holotype: SMF 65628, by original designation. Type locality: "Mingorah, Swat, West-Pakistan". See comment by Stöck, Schmid, Steinlein, and Grosse, 1999, Ital. J. Zool., 66: 230, on type locality, which they considered to be situated at an elevation higher than 1000 m..

Bufo latastii pseudoraddeiHemmer, Schmidtler, and Böhme, 1978, Zool. Abh. Staatl. Mus. Tierkd. Dresden, 34: 377.

Bufo danatensis pseudoraddeiPisanets and Shcherbak, 1979, Vestn. Zool., Kiev, 1979: 11.

Bufo pseudoraddei pseudoraddeiStöck, Schmid, Steinlein, and Grosse, 1999, Ital. J. Zool., 66: 221.

Bufo zamdaensis Fei, Ye, and Huang In Fei, Ye, Huang, and Chen, 1999, Zool. Res., Kunming, 20: 296. Holotype: CIB 760085, by original designation. Type locality: "Zamda (Tzada) County (79° 80′ E, 31° 50′ N), Ngari (Ari) Prefecture, Xizang (Tibet), altitude 2900 m", China. Synonymy by Dufresnes, Mazepa, Jablonski, Oliveira, Wenseleers, Shabanov, Auer, Ernst, Koch, Ramírez-Chaves, Mulder, Simonovo, Tiutenko, Kryvokhyzhar, Wennekes, Zinenko, Korshunov, Al-Johany, Peregontsev, Masroor, Betto-Colliard, Denoël, Borkin, Skorinov, Pasynkova, Mazanaeva, Rosanov, Dubey, and Litvinchuk, 2019, Mol. Phylogenet. Evol., 141: 20. 

Bufo zamdaensis — Fei, Ye, Huang, Jiang, and Xie, 2005, in Fei et al. (eds.), Illust. Key Chinese Amph.: 93.

Pseudepidalea zamdaensis — Frost, Grant, Faivovich, Bain, Haas, Haddad, de Sá, Channing, Wilkinson, Donnellan, Raxworthy, Campbell, Blotto, Moler, Drewes, Nussbaum, Lynch, Green, and Wheeler, 2006, Bull. Am. Mus. Nat. Hist., 297: 365.

Pseudepidalea pseudoraddei — Frost, Grant, Faivovich, Bain, Haas, Haddad, de Sá, Channing, Wilkinson, Donnellan, Raxworthy, Campbell, Blotto, Moler, Drewes, Nussbaum, Lynch, Green, and Wheeler, 2006, Bull. Am. Mus. Nat. Hist., 297: 365.

Bufo (Bufotes) pseudoraddei — Dubois and Bour, 2010, Zootaxa, 2447: 25. See comment under Bufotes record.

Bufo (Bufotes) pseudoraddei pseudoraddei — Dubois and Bour, 2010, Zootaxa, 2447: 25. See comment under Bufotes record.

Bufo (Bufotes) zamdaensis — Dubois and Bour, 2010, Zootaxa, 2447: 25. See comment under Bufotes record.

Bufo pseudoraddei — Kuzmin, 2013, Amph. Former Soviet Union, Ed. 2: 161. 

Bufotes pseudoraddei — Frost, 2013, Amph. Spec. World, Vers. 5.6: . Required change because of non-monophyly of Bufo.

Bufotes pseudoraddei pseudoraddei — Frost, 2013, Amph. Spec. World, Vers. 5.6: . Required change because of non-monophyly of Bufo.

Bufotes zamdaensis — Frost, 2013, Amph. Spec. World, Vers. 5.6. Required change because of non-monophyly of Bufo.

English Names

Swat Green Toad (Bufo pseudoraddei pseudoraddei: Khan, 2002, Bull. Chicago Herpetol. Soc., 37: 159).

Zamda Toad (now longer recognized: Fei, Ye, and Jiang, 2012, Colored Atlas Chinese Amph. Distr.: 267). 

Distribution

Himalayan moist-temperate forest in northern Pakistan, and into nearby Sutlij and Spiti Valley on the western Tibet, China/India border. 

Comment

In the Bufo viridis group Inger, 1972, in Blair (ed.), Evol. Genus Bufo; and the Bufo viridis subgroup of Stöck, Günther, and Böhme, 2001, Zool. Abh. Staatl. Mus. Tierkd. Dresden, 51. Stöck, Schmid, Steinlein, and Grosse, 1999, Ital. J. Zool., 66: 215-232, discussed the subspecific status of Bufo pseudoraddei pseudoraddei and Bufo pseudoraddei baturae (now Bufotes baturae), and considered both to be triploid representatives of the Bufo viridis (sub-) group. Kuzmin, 1999, Amph. Former Soviet Union: 265, previously had considered Bufo viridis pseudoraddei a synonym of the later-described Bufo danatensis. Stöck, Frynta, Grosse, Steinlein, and Schmid, 2001, Asiat. Herpetol. Res., 9: 77-100, showed new records mainly from Pakistan. Stöck, Günther, and Böhme, 2001, Zool. Abh. Staatl. Mus. Tierkd. Dresden, 51: 302-305, provided an account of this triploid species.See preliminary map in Stöck, Günther, and Böhme, 2001, Zool. Abh. Staatl. Mus. Tierkd. Dresden, 51: 308, in connection with Stöck, Frynta, Grosse, Steinlein, and Schmid, 2001, Asiat. Herpetol. Res., 9: App.I. Khan, 2006, Amph. Rept. Pakistan: 46-47, provided a brief account for Pakistan. Stöck, Moritz, Hickerson, Frynta, Dujsebayeva, Eremchenko, Macey, Papenfuss, and Wake, 2006, Mol. Phylogenet. Evol., 41: 663-689, provided molecular evidence of strong differentiation of Bufotes pseudoraddei pseudoraddei and Bufotes pseudoraddei baturae. Ficetola, Crottini, Casiraghi, and Padoa-Schioppa, 2010, North-West. J. Zool., Romania, 6: 1-12, discussed the habitat in Pakistan of Bufotes pseudoraddei baturae (as Bufo). Kuzmin, 2013, Amph. Former Soviet Union, Ed. 2: 161–162, provided an account (as Bufo pseudoraddei) for the former USSR. Dufresnes, Mazepa, Jablonski, Oliveira, Wenseleers, Shabanov, Auer, Ernst, Koch, Ramírez-Chaves, Mulder, Simonovo, Tiutenko, Kryvokhyzhar, Wennekes, Zinenko, Korshunov, Al-Johany, Peregontsev, Masroor, Betto-Colliard, Denoël, Borkin, Skorinov, Pasynkova, Mazanaeva, Rosanov, Dubey, and Litvinchuk, 2019, Mol. Phylogenet. Evol., 141: 17, noted this triploid hybrid-origin species to have originated as Bufotes perrini X Bufotes latastii. A triploid hybrid species (♀ Bufotes perrini (western lineage) × ♂ Bufotes latastii) according to Dufresnes, Mazepa, Jablonski, Oliveira, Wenseleers, Shabanov, Auer, Ernst, Koch, Ramírez-Chaves, Mulder, Simonovo, Tiutenko, Kryvokhyzhar, Wennekes, Zinenko, Korshunov, Al-Johany, Peregontsev, Masroor, Betto-Colliard, Denoël, Borkin, Skorinov, Pasynkova, Mazanaeva, Rosanov, Dubey, and Litvinchuk, 2019, Mol. Phylogenet. Evol., 141: 1–25, who summarized its biogeography and evolutionary history. Che, Jiang, Yan, and Zhang, 2020, Amph. Rept. Tibet: 141–146, provided a detailed account (as Bufotes zamdaensis, in apparent ignorance of the recent 2019 synonymy with Bufotes pseudoraddei) for Tibet

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